{"key":"f0803c8a-f55c-4f22-9004-3c4cb4a535b5","installationKey":"68845aab-a453-4816-b36d-a3ec8455deb0","publishingOrganizationKey":"2c39be5c-c11e-46d0-bcb4-552f2072d19f","doi":"10.15472/m5ewf1","version":"1.1","external":false,"numConstituents":0,"type":"CHECKLIST","subtype":"TAXONOMIC_AUTHORITY","shortName":"Scarabaeinae de Colombia","title":"Listado de las especies de la subfamilia Scarabaeinae (Coleoptera: Scarabaeidae) de Colombia","description":"<p>Este recurso presenta un listado actualizado de las especies de escarabajos de la subfamilia Scarabaeinae (Coleoptera: Scarabaeoidea: Scarabaeidae) reportadas en la literatura para el territorio colombiano. El listado incluye 10 tribus, 40 géneros, 351 especies y 7 subespecies. Adicionalmente, se identifican una tribu, un género y seis especies como sinónimos, cuyos nombres aún son citados en la literatura, en algunos casos debido a su uso histórico o a la falta de adopción de actualizaciones taxonómicas recientes. Se reconocen 65 especies y una subespecie como presumiblemente endémicas de Colombia, dado que su distribución geográfica ha sido registrada exclusivamente en el país. El único taxón introducido corresponde al género Digitonthophagus, representado por la especie Digitonthophagus gazella. En cuanto al estado de amenaza, dos especies están categorizadas como En Peligro (EN), tres como Casi Amenazadas (NT), 34 como de Preocupación Menor (LC), 13 como Datos Deficientes (DD) y 308 no han sido evaluadas (NE). El listado documenta la distribución geográfica a nivel de países y departamentos dentro de Colombia, el medio de establecimiento, el estatus taxonómico, el estado de amenaza, los nombres originales y otra información relevante para cada especie.</p><p> La compilación se basa en los principales catálogos especializados del grupo, literatura taxonómica reciente, descripciones originales, revisiones taxonómicas, información sobre ejemplares tipo y perfiles de especie. El conjunto de datos incluye 445 referencias bibliográficas asociadas a los taxones, que documentan la fauna de Scarabaeinae de Colombia. Este número puede diferir del total de referencias citadas en el metadato del recurso, el cual incorpora únicamente las fuentes principales empleadas para la compilación y contextualización del listado. Para algunas especies, la presencia en Colombia no ha sido corroborada mediante revisiones taxonómicas recientes o evidencia adicional; dichas situaciones se documentan explícitamente en el campo taxonRemarks de la lista y en el campo description de la hoja de referencias del conjunto de datos.</p>","language":"spa","logoUrl":"https://gitlab.com/sib-colombia/logos/-/raw/main/socio-SiB-coleocol.png","citation":{"text":"Martínez-Revelo D E, Medina-Uribe C A, González-Alvarado A, Chamorro-Fuertes W R, Arias-Buriticá J A (2026). Listado de las especies de la subfamilia Scarabaeinae (Coleoptera: Scarabaeidae) de Colombia. Version 1.1. Coleoptera de Colombia. 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JC, Pérez-Buitrago N, Lopera A (2023) Registros de escarabajos coprófagos (Coleoptera: Scarabaeidae) depositados en la Colección Entomológica de la Orinoquia (CEO). 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Revista Colombiana de Entomología 45(2): 1-10 e7963","identifier":"https://doi.org/10.25100/socolen.v45i2.7963","citationProvidedBySource":false},{"text":"Amézquita-Melo S, Forsyth A, Lopera-Toro A, Camacho A (1999) Comparación de la composición y riqueza de especies de escarabajos coprófagos (Coleoptera: Scarabaeidae) en remanentes de bosque de la Orinoquía Colombiana Acta Zoológica Mexicana (n. s.) 76:113-126","identifier":"https://www.redalyc.org/pdf/575/57507608.pdf","citationProvidedBySource":false},{"text":"Arango L, Montes J (2009) Caracterización entomológica parcial de la cuenca del río La Miel en el departamento de Caldas. Boletín científico del museo de historia natural. 13 (2): 249 - 268","identifier":"http://www.scielo.org.co/scielo.php?script=sci_arttext&pid=S0123-30682009000200016","citationProvidedBySource":false},{"text":"Arias-Buriticá JA (2011) Revisión taxonómica de la Sección “Buqueti”, Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae). 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El único taxón introducido corresponde al género Digitonthophagus, representado en el país por Digitonthophagus gazella; las demás especies incluidas en el conjunto de datos se consideran nativas de los ecosistemas colombianos.","coverages":[{"scientificName":"Scarabaeidae","rank":{"verbatim":"family","interpreted":"FAMILY"}},{"scientificName":"Scarabaeinae","rank":{"verbatim":"subfamily","interpreted":"SUBFAMILY"}},{"scientificName":"Ateuchini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Canthonini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Coprini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Deltochilini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Dichotomiini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Eurysternini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Homocoprini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Oniticellini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Onthophagini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Phanaeini","rank":{"verbatim":"tribe","interpreted":"TRIBE"}},{"scientificName":"Agamopus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Andinocopris","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Anisocanthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Anomiopus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Ateuchus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Bdelyrus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Canthidium","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Canthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Canthonella","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Copris","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Coprophanaeus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Cryptocanthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Deltochilum","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Dendropaemon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Diabroctis","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Dichotomius","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Digitonthophagus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Eurysternus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Genieridium","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Gromphas","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Hansreia","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Homocopris","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Malagoniella","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Ontherus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Onthophagus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Oruscatus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Oxysternon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Phanaeus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Pseudocanthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Scatimus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Scatonomus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Scybalocanthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Sinapisoma","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Sulcophanaeus","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Sylvicanthon","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Tetraechma","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Tetramereia","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Trichillidium","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Trichillum","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Uroxys","rank":{"verbatim":"genus","interpreted":"GENUS"}},{"scientificName":"Canthidium (Canthidium)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthidium (Neocanthidium)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthon (Canthon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthon (Francmonrosia)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthon (Glaphyrocanthon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthon (Goniocanthon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Canthon (Pseudepilissus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Coprophanaeus (Coprophanaeus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Coprophanaeus (Megaphanaeus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Deltochilum (Calhyboma)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Deltochilum (Deltochilum)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Deltochilum (Deltohyboma)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Deltochilum (Euhyboma)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Deltochilum (Hybomidium)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dendropaemon (Crassipaemon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dendropaemon (Dendropaemon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dendropaemon (Glaphyropaemon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dendropaemon (Nigropaemon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dendropaemon (Rutilopaemon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dichotomius (Cephagonus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dichotomius (Dichotomius)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dichotomius (Luederwaldtius)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Dichotomius (Selenocopris)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Malagoniella (Malagoniella)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Ontherus (Caelontherus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Ontherus (Ontherus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Onthophagus (Onthophagus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Oxysternon (Mioxysternon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Oxysternon (Oxysternon)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}},{"scientificName":"Phanaeus (Phanaeus)","rank":{"verbatim":"subgenus","interpreted":"SUBGENUS"}}]}],"geographicCoverages":[{"description":"Se reportan especies de la subfamilia Scarabaeinae (Coleoptera: Scarabaeoidea: Scarabaeidae) en todo el territorio nacional. Se especifica la distribución a nivel de países y departamentos al interior de Colombia. Además, se incluyen especies sin localidad específica y algunos registros departamentales en literatura que deben ser verificados o actualizados","boundingBox":{"minLatitude":-90.0,"maxLatitude":90.0,"minLongitude":-180.0,"maxLongitude":180.0,"globalCoverage":true}}],"temporalCoverages":[{"@type":"range","start":"1758-01-01T00:00:00.000+00:00","end":"2025-12-31T00:00:00.000+00:00"}],"keywordCollections":[{"thesaurus":"SiB Colombia","keywords":["Lista de referencia"]},{"thesaurus":"N/A","keywords":["Sur América","géneros","especies","escarabajos estercoleros","inventario","literatura"]},{"thesaurus":"GBIF Dataset Type Vocabulary: http://rs.gbif.org/vocabulary/gbif/dataset_type_2015-07-10.xml","keywords":["Checklist"]},{"thesaurus":"GBIF Dataset Subtype Vocabulary: http://rs.gbif.org/vocabulary/gbif/dataset_subtype.xml","keywords":["Taxonomicauthority"]}],"project":{"title":"Listado de los coleópteros de Colombia","contacts":[{"type":"POINT_OF_CONTACT","primary":false,"userId":["https://orcid.org/0000-0002-4446-4902"],"firstName":"Diego Esteban","lastName":"Martínez-Revelo","position":[],"email":[],"phone":[],"homepage":[],"address":[]}],"funding":"Fondos para la estancia de DEMR en la colección Entomológica del Instituto Humboldt (IAvH-E), Villa de Leyva, Boyacá, fueron obtenidos del proyecto “Impacts of tropical agriculture on biodiversity across spatial scales” financiado por el “Research Council of Norway\" (Grant/Award Number: 262378) y realizado por la “Norwegian University of Life Sciences” y el Instituto Alexander von Humboldt (Convenio de Cooperación No. 11-256).","awards":[],"studyAreaDescription":"Se compilan los nombres de las especies de Coleoptera que se han registrado para Colombia de acuerdo con la literatura.","designDescription":"Toda la información presentada en estos listados proviene de la literatura: catálogos, listados previos, revisiones taxonómicas y otros reportes.","relatedProjects":[],"abstract":"El listado de los coleópteros de Colombia es una iniciativa del Grupo de Coleopterólogos de Colombia (ColeopCol, https://sites.google.com/view/coleopcol), que pretende compilar el listado de las especies de coleópteros registradas para Colombia en la literatura. El proyecto no está adscrito a ninguna institución, sino que nace del acuerdo entre coleopterólogos colombianos al ver la necesidad de compilar, organizar y difundir esta información"},"samplingDescription":{"studyExtent":"El listado compila los nombres de las especies de la subfamilia Scarabaeinae (Coleoptera: Scarabaeoidea: Scarabaeidae) mencionados en literatura para el territorio colombiano.","sampling":"La información del recurso se compiló a partir de una revisión exhaustiva de la literatura taxonómica y faunística disponible sobre la subfamilia Scarabaeinae (Coleoptera: Scarabaeoidea: Scarabaeidae) para el territorio colombiano. Se recopilaron los nombres científicos válidos, autores, sinonimias relevantes y referencias bibliográficas de las especies registradas en el país.\n\nLa lista de géneros se fundamentó en publicaciones nacionales de referencia (Contreras 1951; Escobar 2000; Medina et al. 2001; Noriega et al. 2015) y en trabajos de alcance continental para las Américas (Vaz-de-Mello et al. 2011). Para las especies a nivel nacional, se consultaron publicaciones históricas y recientes, incluyendo catálogos, listas de especies y revisiones taxonómicas (Gillet 1911; Blackwelder 1945; Contreras 1951; Escobar 2000; Medina et al. 2001; Krajcik 2012; Chamorro et al. 2020).\n\nAdicionalmente, se revisaron listados regionales y departamentales (p. ej., Sierra Nevada de Santa Marta, Orinoquia, Andes, Caribe colombiano, bosque seco tropical y Eje Cafetero), así como revisiones taxonómicas de géneros y grupos de especies que incluyeron material colombiano o especies endémicas del país.\n\nCon base en la distribución geográfica reportada en la literatura, se asignaron categorías biogeográficas a las especies (endémicas, nativas o introducidas), siguiendo a Chamorro et al. (2020)y Noriega et al. (2020). Asimismo, se consultó la Lista Roja de la UICN para clasificar las especies según su categoría de amenaza (EN, VU, NT, LC, DD, NE).\n\nFinalmente, se consultó la World Scarabaeidae Database (versión 2023-09-05) para verificar la información nomenclatural y los datos de los tipos de las especies incluidas, incorporando actualizaciones derivadas de revisiones taxonómicas recientes.","qualityControl":"Los nombres científicos incluidos en el listado fueron verificados mediante la consulta de las descripciones originales, catálogos taxonómicos y revisiones recientes citadas en el recurso. Se revisó la validez nomenclatural de los nombres, su autoría, año de descripción y estatus taxonómico actual, así como la correcta asignación a género, subgénero y tribu. Adicionalmente, se evaluó la consistencia interna del listado para evitar duplicaciones derivadas de sinonimias o cambios nomenclaturales, y se contrastó la información con bases de datos taxonómicas especializadas cuando fue necesario.","methodSteps":["El levantamiento de las especies de la subfamilia Scarabaeinae registradas para Colombia se realizó mediante una compilación sistemática de información bibliográfica. Como punto de partida se utilizaron catálogos y listas de referencia para el país y la región, incluyendo a Blackwelder (1945), Contreras (1951), Escobar (2000) y Medina et al. (2001). A partir de estas fuentes base, se revisaron publicaciones taxonómicas y faunísticas más recientes para incorporar las especies descritas o registradas para Colombia hasta la fecha de elaboración del conjunto de datos.","Para cada especie, se registró el nombre científico, autoría, año de descripción y referencias bibliográficas asociadas. La grafía, autoría y estatus nomenclatural de los nombres fueron verificados mediante la consulta de las descripciones originales, catálogos taxonómicos y revisiones recientes citadas en el recurso. Adicionalmente, se utilizó la herramienta Species Matching de GBIF como apoyo para la detección de variantes ortográficas, sinonimias y posibles inconsistencias nomenclaturales, sin sustituir la verificación basada en literatura especializada.","Las referencias bibliográficas fueron compiladas a partir de descripciones originales, nuevos registros para el país, revisiones taxonómicas y actos nomenclaturales (por ejemplo, sinonimias, revalidaciones y cambios de estatus). Cuando existieron discrepancias entre fuentes, se priorizaron las revisiones taxonómicas más recientes y aquellas que incluyeron material colombiano.","La información final fue organizada en una plantilla estandarizada, asegurando la consistencia taxonómica entre géneros, subgéneros, tribus y especies, así como la correspondencia entre los nombres válidos y sus referencias. Este procedimiento permite que el conjunto de datos pueda ser interpretado y reproducido por otros investigadores a partir de las mismas fuentes bibliográficas."]},"countryCoverage":[],"collections":[{"name":"Canadian Museum of Nature, Insect Collection (CMNEN | HAHC)","identifier":"https://www.gbif.org/grscicoll/collection/26e0367f-d400-46c5-b37f-978a5b229c07","curatorialUnits":[]},{"name":"Colección Alejandro Lopera Toro-Escarabajos Coprófagos de Colombia","identifier":"https://www.gbif.org/grscicoll/collection/e6def1ec-3d01-4362-9e53-ae6d77c977ad","curatorialUnits":[]},{"name":"Colección de Zoología del Instituto de Ciencias Naturales de la Universidad Nacional de Colombia","identifier":"https://www.gbif.org/grscicoll/collection/32b51f25-9fb2-4086-8631-0b60a0adca1c","curatorialUnits":[]},{"name":"Colección Entomológica de Mato Grosso Eurides Furtado","identifier":"https://www.gbif.org/grscicoll/institution/4bbb3e94-174e-4866-a7c4-3c2e7ba60d97","curatorialUnits":[]},{"name":"Colecciones Biológicas del Instituto Alexander von Humboldt","identifier":"https://www.gbif.org/grscicoll/collection/f707aa14-d87b-4fec-a081-28f3e4ec6a6f","curatorialUnits":[]},{"name":"Instituto Nacional de Biodiversidad (INBio)","identifier":"https://www.gbif.org/grscicoll/institution/bcd366c3-0996-47b5-9b9b-ff72bba38e78","curatorialUnits":[]},{"name":"Museo Argentino de Ciencias Naturales Bernardino Rivadavia (MACN)","identifier":"https://www.gbif.org/grscicoll/institution/8c4ae816-dfd9-4bc8-a2fd-f25a56bf82e9","curatorialUnits":[]},{"name":"Museo Zoologico \"La Specola\", Italy, Firenze","identifier":"https://www.gbif.org/grscicoll/institution/6df7cd8a-3b8f-4cb3-a6ae-fe3d7398f0e7","curatorialUnits":[]},{"name":"Museu de Zoologia da Universidade de Sao Paulo (MZSP)","identifier":"https://www.gbif.org/grscicoll/institution/571a17f5-b06d-4fc8-8c74-a18ff480b304","curatorialUnits":[]},{"name":"Muséum national d’Histoire naturelle (MNHN)","identifier":"https://www.gbif.org/grscicoll/institution/6a6ac6c5-1b8a-48db-91a2-f8661274ff80","curatorialUnits":[]},{"name":"National Museum Prague","identifier":"https://www.gbif.org/grscicoll/institution/1eb81ba8-4c20-4637-b120-ce3e262b83e6","curatorialUnits":[]},{"name":"Royal Belgian Institute of Natural Sciences (RBINS)","identifier":"https://www.gbif.org/grscicoll/institution/c2bfdeef-9c03-435e-8465-c483dadd6995","curatorialUnits":[]},{"name":"Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute","identifier":"https://www.gbif.org/grscicoll/institution/f444dc20-d121-4462-8ea0-2a53df1444ad","curatorialUnits":[]},{"name":"Sweden, Stockholm, Naturhistoriska riksmuseet","identifier":"https://www.gbif.org/grscicoll/institution/5ed4ae5a-4638-4c61-8af1-b476f64bc0a2","curatorialUnits":[]},{"name":"The Natural History Museum, London, United Kingdom","identifier":"https://www.gbif.org/grscicoll/institution/1d808a7c-1f9e-4379-9616-edb749ecf10e","curatorialUnits":[]},{"name":"Zoologische Staatssammlung München (snsb)","identifier":"https://www.gbif.org/grscicoll/institution/774f65a5-ed87-4356-9515-ea9fd8d68650","curatorialUnits":[]}],"dataDescriptions":[{"name":"Listados de especies publicados por el Grupo de Coleopterólogos","charset":"UTF-8","url":"https://sites.google.com/view/coleopcol/listados","format":"Página web","formatVersion":"1"}],"dataLanguage":"spa","pubDate":"2026-02-18T00:00:00.000+00:00","maintenanceUpdateFrequency":"AS_NEEDED","maintenanceDescription":"","license":"http://creativecommons.org/licenses/by/4.0/legalcode","dwca":{"coreType":"http://rs.tdwg.org/dwc/terms/Taxon","extensions":["http://rs.gbif.org/terms/1.0/SpeciesProfile","http://rs.gbif.org/terms/1.0/TypesAndSpecimen","http://rs.gbif.org/terms/1.0/Reference","http://rs.gbif.org/terms/1.0/Distribution"],"modified":"2026-10-19T01:56:15.592+00:00"},"approximateCounts":{"occurrenceCount":0,"nameUsageCount":593}}