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It is a member of the genus Notholaena, which is part of the subfamily Cheilanthoideae of family Pteridaceae. It is distinguished by the pentagonal shape formed by the blades of its frond, a property other members of Notholaena do not possess."},{"description":"Notholaena standleyi is indigenous to southwestern United States and parts of north and central Mexico. It specifically grows in Arizona, Colorado, New Mexico, Oklahoma, and Texas in the USA. Map of Notholaena standleyi distribution in the US"},{"description":"Notholaena standleyi is a perennial species that typically grows in desert regions at elevations from 300 to 2100 m. It is found on rocky hillsides, usually in the crevices created by limestone and granite boulders that provide the partial shade the plant prefers. During periods of drought, the frond may curl and become brown until water is available, an adaptation to the semi-arid environments it inhabits. At lower elevations, it sometimes grows alongside Notholaena californica. It takes 13 days to germinate and sporulates from late spring to fall. It has the same life cycle as a typical fern, the sporophyte produces haploid spores by meiosis, which grows into a haploid gametophyte. The gametophyte produce gametes by mitosis. The fertilized egg when develop into the sporophyte by mitosis. The spore is transported by air and water."},{"description":"Individuals of this species have brown petioles and green fronds. The ferns are erect and grow to a height of 25 cm. The petioles grow to a length of 5 to. The individual fronds are usually 3 to wide as well as long. The frond is light green when young, becoming darker green as it matures, and usually contains five blades arranged in a pentagonal fashion. The blades are bipinnately or tripinnately compound, and each of them is triangular and pointed. The abaxial side of the frond is covered by powdery farina, with a function that is still unclear. The color of the farina differs between species growing in different geographic locations. These are referred to as chemotypes, of which there are four. These chemotypes also have different number of complete chromosome sets. Gold and yellow farina are diploids, pallid and yellow/green are tetraploids. Its prothallus contains glandular trichomes. It usually has 32 spores per sporangium, but many with only 16 have been observed, produced from eight 2n mother cells. It is diploid and has 60 chromosomes in its root tip cells."},{"description":"Notholaena standleyi was named after Paul C. Standley, Assistant Curator of the United States National Herbarium, who studied plants in southwestern United States. First described as Notholaena candida var. quinquefidopalmata in 1864, it was elevated to a species as Notholaena hookeri in 1879 by D. C. Eaton. However, this name had already been used to describe another fern, and it was renamed by William R. Maxon in 1915. He named it after Standley, who had engaged in a great deal of field study in the southwestern United States. Standley was also collaborating with E. O. Wooton to develop a comprehensive Flora of New Mexico at the time."},{"description":"The Seri Indians made tea from the fronds to promote fertility in women. The species was also believed to have supernatural value; the Seri carried in a small cloth bag to bring good luck to the bearer. It has been studied in potential herpes treatments, because of its antiviral activity. Notholaenic acid, isolated from the species, has shown anti-HSV-1 activity at high concentrations (25 μg/disk)."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"165713018":"Notholaena"}},{"key":207631335,"datasetKey":"f382f0ce-323a-4091-bb9f-add557f3a9a2","nubKey":2651608,"parentKey":1026675,"parent":"Hemionitis","acceptedKey":311266907,"accepted":"Hemionitis standleyi (Maxon)","family":"Pteridaceae","genus":"Hemionitis","species":"Hemionitis standleyi","familyKey":322132490,"genusKey":207630951,"speciesKey":311266907,"scientificName":"Notholaena standleyi Maxon","canonicalName":"Notholaena standleyi","authorship":"Maxon","publishedIn":"Amer. 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It has leathery, thrice-divided leaves with dark brown axes; the leaves are coated with white powder below. First described as a species in 1853, it was transferred to the new genus Argyrochosma (the \"false cloak ferns\") in 1987, recognizing their distinctness from the \"cloak ferns\" (Notholaena sensu stricto)."},{"description":"The horticulturist George Schneider considered the species suitable for greenhouse cultivation."},{"description":"Argyrochosma chilensis is a small epipetric fern. The rhizome is stout and upright. It bears thin, linear scales 3 mm long that terminate in a hair attached by a joint, of a bright brown color, with entire (toothless) margins. From it, the fronds arise in clumps. From base to tip of leaf, they are 5 to long. Of this length, about half is made up by the stipe (the stalk of the leaf, below the blade), which is round, a dark chestnut brown in color, and lacks scales and hairs. The leaf blades are ovate, broadest at the lowest or second pair of pinnae, with an acute (pointed) tip. They are tripinnate (cut into pinnae, pinnules, and pinnulets), typically bearing about five pairs of pinnae. The texture of the blade is leathery; the underside is densely covered in white farina (powder), which is absent from the upper side. The pinnae are ovate-lanceolate to ovate, attached to the rachis (leaf axis) by short stalks. Each pinna is tipped with a three-lobed, rhomboid (diamond-shaped) terminal pinnule; the other pinnules are cut into pinnulets, which are oblong, obtuse (blunt) at the tip, and are generally attached directly to the costule (pinnule axis), without stalks. They are not jointed at the base. The sori lie along the veins, in the one-third to one-fourth of the vein closest to the leaf edge. The leaf edges are curved under, but not otherwise modified into false indusia. Each sporangium contains 64 spores. The spores are covered with a network of raised crests, although portions of the surface lack elevated crests."},{"description":"Argyrochosma chilensis is endemic to the Juan Fernández Islands, particularly Robinson Crusoe Island and Alejandro Selkirk Island. It is found on rocks in dry, exposed situations, at altitudes from 20 to."},{"description":"Notes References Works cited"},{"description":"The first scientist to mention the species was Antoine Laurent Apollinaire Fée in 1852, who referred to it as Pellaea chilensis without offering a species description (making that name a nomen nudum), but noted that Jules Rémy, who was preparing a flora of Chile, considered it to belong to Lindsaea, in the sense used by Desvaux. Rémy published his flora the following year, in the sixth botanical volume of Claude Gay's Historia fisica y politica de Chile, and described the species as Cincinalis chilensis, attributing that name to Fée. While he did not explain his choice of epithet, it presumably reflects the Chilean origin of the specimens he examined. In 1858, Johann Wilhelm Sturm transferred it into a broadly-construed genus Notholaena as N. chilensis. Carl Christensen, by contrast, assigned it to Pellaea as P. chilensis in his Index Filicum of 1906. In 1856, Edward Joseph Lowe described a fern being cultivated in Britain under the name of Notholaena nivea which, however, had a broader blade, more heavily dissected and with more leaf tissue, than the leaves of true N. nivea. He was unable to find a corresponding specimen in the herbarium at Kew, and described it as a new species, which he called Notholaena hookeri, in honor of William Jackson Hooker. He attributed its introduction to British cultivation to John Riley, of Papplewick. In 1866, John Smith transferred it to Cincinalis as C. hookeri. The name \"N. hookeri\" was mistakenly reused by Daniel Cady Eaton; William Ralph Maxon, when devising the replacement name of Notholaena standleyi for Eaton's fern in 1915, opined that N. hookeri, as named by Eaton, was simply a synonym of N. nivea. Rolla M. Tryon Jr. designated a specimen of Riley's cultivation at Kew as the lectotype, and concluded that it was in fact synonymous with N. chilensis. While Tryon, when finishing Charles Alfred Weatherby's revision of American Notholaena, considered it impossible to reasonably subdivide Notholaena into sections based on the data available at the time, both Edwin Copeland and Weatherby himself had suggested in the 1940s that a group of ferns related to N. nivea might represent a distinct genus. This was finally addressed in 1987 by Michael D. Windham, who was carrying out phylogenetic studies of these genera. He elevated Notholaena sect. Argyrochosma to become the genus Argyrochosma, and transferred this species to that genus as A. chilensis. In 2018, Maarten J. M. Christenhusz transferred the species to Hemionitis as H. chilensis, as part of a program to consolidate the cheilanthoid ferns into that genus. He also transferred N. hookeri, as H. hookeri. A phylogenetic analysis including a single specimen of A. chilensis found it nested within a clade representing A. nivea sensu lato. It probably evolved by the dispersal of A. nivea to Robinson Crusoe Island, followed by anagenesis creating A. chilensis and its spread to the younger Alejandro Selkirk Island."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"165621828":"Argyrochosma"}},{"key":186750140,"datasetKey":"bae5856f-da10-4333-90a0-5a2135361b30","nubKey":2651610,"parentKey":983302,"parent":"Notholaena","acceptedKey":211424778,"accepted":"Notholaena standleyi Maxon","order":"Polypodiales","family":"Pteridaceae","genus":"Notholaena","species":"Notholaena standleyi","orderKey":186716297,"familyKey":186745461,"genusKey":211424773,"speciesKey":211424778,"scientificName":"Cheilanthes hookeri (Kümmerle) Domin","canonicalName":"Cheilanthes hookeri","authorship":"(Kümmerle) Domin","nameType":"SCIENTIFIC","taxonomicStatus":"SYNONYM","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"Cheilanthes_hookeri_Kümmerle_Domin","habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":true,"higherClassificationMap":{"186716297":"Polypodiales","186745461":"Pteridaceae","211424773":"Notholaena","211424778":"Notholaena standleyi","983302":"Notholaena"}},{"key":301861560,"datasetKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":3750206,"parentKey":1944164,"parent":"Notholaena","acceptedKey":301861556,"accepted":"Notholaena standleyi (Kümmerle) Maxon","kingdom":"Plantae","phylum":"Tracheophyta","order":"Polypodiales","family":"Pteridaceae","genus":"Notholaena","species":"Notholaena standleyi","kingdomKey":293994071,"phylumKey":294954977,"classKey":296010258,"orderKey":301826245,"familyKey":301860465,"genusKey":301861547,"speciesKey":301861556,"scientificName":"Chrysochosma hookeri (Domin) Kümmerle","canonicalName":"Chrysochosma hookeri","authorship":"(Domin) Kümmerle","publishedIn":"Kümmerle. 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