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N (colonies, zooids) Mean SD Range Zooid length 3, 30 414 39 367 – 489 Zooid width 3, 30 333 32 294 – 384 Orifice length 3, 15 100 13 82 – 116 Orifice width 3, 15 93 3 90 – 95 Ovicell length 2, 3 180 3 177 – 182 Ovicell width 2, 3 219 11 206 – 228 N, Number of colonies and number of zooids measured; SD, standard deviation."},{"description":"Remarks. None of the material examined of this species had an intact ovicell; in all instances the roof of the ovicell was broken, revealing a floor consisting of an exterior skeletal wall with planar spherulitic microstructure delineating two lobes on either side of a median depression (Fig. 41). The presence of a small kenozooid distal of each ovicell, which is unique among the species described in this paper, recalls various other cheilostomes (e. g. Bishop & Househam 1987; Ostrovsky et al. 2008), and may signify evolutionary reduction of the zooid distal of the maternal zooid from an autozooid to a kenozooid. Hayward & Ryland (1993) introduced the new genus Apiophragma for M. hyalina Waters, 1904, referring to the shape of the mural rim from the Greek apios, a pear and phragma, a wall. Apiophragma resembles Megapora in its depressed cryptocyst, thick mural rim and distal arc of spines but differs in having only a weakly trifoliate opesia which is much smaller and narrowly bell-shaped, and the presence of opesiules. The inclusion of Apiophragma in the new family Pyrisinellidae should be considered after restudy of the type material of A. hyalina."},{"description":"Description. Colony encrusting, multiserial, unilaminar, developing small rounded patches. Pore chambers present, distal pore chamber slightly larger than distolateral pore chambers; pore windows oval (mean L = 37 µm, mean W = 10 µm), surrounded by a mural ring, that of the distal pore chamber facing frontally. Ancestrula oval and smaller than astogenetically mature autozooids (L = 223 µm, mean W = 166 µm); gymnocyst absent or hidden by later zooids, cryptocyst densely granular, occupying half of the frontal area, opesia trifoliate with two small oral spines (D = 12 µm) placed distally (Fig. 44). Autozooids in zone of early astogeny transitional in size and morphology between ancestrula and later autozooids (Fig. 43). Autozooids small, rounded hexagonal or oval, longer than broad (mean L / W = 1.27), separated by deep furrows. Gymnocyst narrow, slightly broader proximally. Cryptocyst extensive, coarsely and densely granular, depressed centrally, submarginally raised to form a rounded ridge which together with distal rim of the opesia forms a pear-shaped outline on the frontal wall and around the opesia. Opesia strongly trifoliate, the proximal edge gently convex with rounded indentations at the proximolateral corners; a pair of rounded thick lateral denticles directed downwardly divide the larger, distal semicircular part of the opesia from the smaller but broader proximal part (Fig. 41). Oral spines numbering six, arranged in an arch around the distal edge of the opesia, the most proximal pair almost level with the denticles; distalmost pair of oral spines spaced more widely in ovicellate than non-ovicellate zooids. Complete ovicell not observed but that figured by Hayward & Ryland (1998, p. 197, fig. 58 B) is hyperstomial, prominent, rounded and elongated, with a narrow, median triangular ectooecial window with thickened edges; a small rounded triangular kenozooid with a cryptocystal frontal area and a tiny central lacuna is present immediately distally of each ovicell (Fig. 41). Closure plates developed in some autozooids as distal extensions of the cryptocyst, with a small tuberculum located medially level with the proximal edge of the opesia, which is completely occluded (Fig. 40). Kenozooids of irregular shape frequently present along lateral margins of lobate colonies, slightly smaller (L = 258 − 291 µm, W = 195 − 218 µm) than autozooids; gymnocyst narrow but wider than in autozooids; cryptocyst coarsely and densely granular, continuous over the entire frontal area except for a small, round, lacuna (mean D = 25 µm) near the centre (Fig. 42). Avicularia absent."},{"description":"Megapora ringens is an encrusting bryozoan, present in the north of the British Isles. The colonies develop as small inconspicuous round patches. The species occurs in subtidal waters between 150 – 300 m, where it most frequently colonises stones. Megapora ringens is widely distributed in the northeast Atlantic, where it ranges from the Arctic south to Bergen (Norway) and the Shetland Isles."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","53":"Bryozoa","252":"Gymnolaemata","467":"Cheilostomatida","5797":"Calloporidae","1007128":"Megapora"},"class":"Gymnolaemata"},{"key":1007364,"nameKey":10423986,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1007364,"parentKey":1007358,"parent":"Siphonicytara","kingdom":"Animalia","phylum":"Bryozoa","order":"Cheilostomatida","family":"Siphonicytaridae","genus":"Siphonicytara","species":"Siphonicytara hexaserialis","kingdomKey":1,"phylumKey":53,"classKey":252,"orderKey":467,"familyKey":6022,"genusKey":1007358,"speciesKey":1007364,"scientificName":"Siphonicytara hexaserialis Guha & Gopikrishna, 2007","canonicalName":"Siphonicytara hexaserialis","authorship":"Guha & Gopikrishna, 2007","publishedIn":"Guha, A. 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K., & Gopikrishna, K. (2007). Some fossil ascophoran bryozoan taxa from Tertiary sequences of western Kachchh, Gujarat. 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Early Miocene: Spain, Corsica, Italy, Malta (Moissette 1996). Middle Miocene: Austria, Hungary (Moissette et al. 2006), Czech Republic (Zágoršek 2010 b), northern Italy, Calabria, Malta (Moissette 1996). Late Miocene: Algeria (Moissette 1988), Sardinia, Sicily, Calabria (Pizzaferri & Braga 2000), Malta (PM, pers. obs.), Crete (Moissette et al. 1993). Pliocene: Spain, Algeria (Haddadi-Hamdane 1996), northern Italy, Sicily, Crete (PM, pers. obs.), Karpathos (PM, pers. obs.). Pleistocene: Rhodes (PM, pers. obs.). This fossil species has exclusively been recorded from the Mediterranean-Paratethys realm (Moissette 1996). Three present-day Batopora species are known from the Indo-Pacific at depths between 285 and 880 m (Cook & Lagaaij 1976; Hayward & Cook 1979)."},{"description":"REMARKS A few juvenile colonies are present in a fair number of samples. This was also observed by several authors (Cook & Lagaaij 1976; Pizzaferri & Braga 2000)."},{"description":"DESCRIPTION Small conical conescharelliniform colonies with a flattened base an apical tube comprised of kenozooids and terminated by a small pit. Hexagonal zooids arranged in concentric alternating series. Frontal convex with fairly large pores. Large circular aperture located in the distal part of each zooid. No avicularia. Rare small broken hyperstomial ovicells are visible (Fig. 12 A)."},{"description":"(Fig. 12 A-G)"}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","53":"Bryozoa","252":"Gymnolaemata","467":"Cheilostomatida","2097":"Batoporidae","1007438":"Batopora"},"class":"Gymnolaemata"},{"key":1007498,"nameKey":10679527,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1007498,"parentKey":1007486,"parent":"Steginoporella","kingdom":"Animalia","phylum":"Bryozoa","order":"Cheilostomatida","family":"Steginoporellidae","genus":"Steginoporella","species":"Steginoporella vicksburgica","kingdomKey":1,"phylumKey":53,"classKey":252,"orderKey":467,"familyKey":6023,"genusKey":1007486,"speciesKey":1007498,"scientificName":"Steginoporella vicksburgica Canu & Bassler, 1920","canonicalName":"Steginoporella vicksburgica","authorship":"Canu & Bassler, 1920","publishedIn":"Pouyet, S., & David, L. (1979). 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(2021). Seek and ye shall find: new species and new records of Microporella (Bryozoa, Cheilostomatida) in the Mediterranean. ZooKeys, 1053: 1–42. https://doi.org/10.3897/zookeys.1053.65324","nameType":"SCIENTIFIC","taxonomicStatus":"SYNONYM","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1009073","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"The species occurs in subtidal waters"},{"description":"Diporula verrucosa is a warm temperate species that occurs only rarely in the British Isles. It is found more commonly from the coasts of West Africa throughout the Mediterranean. In the UK the species has been recorded off the south Cornish coast where it is as the extreme northern limit of its range."},{"description":"Diporula verrucosa is an erect bryozoan. Colonies arise from an encrusting base, developing into rigid, erect flat fan-shaped structures composed of irregular cylindrical branches. Colonies are light brownish in colour. Autozooids are broad, hexagonal to oval in shape. They are 0.6-0.7 by 0.5-0.6 mm."}],"vernacularNames":[],"synonym":true,"higherClassificationMap":{"1":"Animalia","53":"Bryozoa","252":"Gymnolaemata","467":"Cheilostomatida","5826":"Microporellidae","1008982":"Microporella","11300007":"Microporella verrucosa"},"class":"Gymnolaemata"},{"key":1009224,"nameKey":3110325,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1009224,"parentKey":1009223,"parent":"Cupuladria","kingdom":"Animalia","phylum":"Bryozoa","order":"Cheilostomatida","family":"Cupuladriidae","genus":"Cupuladria","species":"Cupuladria biporosa","kingdomKey":1,"phylumKey":53,"classKey":252,"orderKey":467,"familyKey":5803,"genusKey":1009223,"speciesKey":1009224,"scientificName":"Cupuladria biporosa (Canu & Bassler, 1923)","canonicalName":"Cupuladria biporosa","authorship":"(Canu & Bassler, 1923) ","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1009224","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Material examined. VMNH no. 70611, 70612; USNM no. 1283236."},{"description":"Distribution. Cape Hatteras to Brazil, Caribbean, Gulf of Mexico."},{"description":"Remarks. Three cupuladriid species are common along the continental shelf from Cape Hatteras to Florida according to the study of Maturo (1968). The two smaller species, Cupuladria doma and Discoporella depressa, occur across the entire shelf, but Cupuladria biporosa was found only on the outer part of the shelf. Their occurrence in Oculina rubble may represent a death assemblage as no colonies found had been alive when collected, but sediment samples from the Florida shelf generally contain a large number of dead, relative to living, colonies of cupuladriids, so the species may well be part of the Oculina reef community."},{"description":"Description. Saucer-shaped colonies, free-living on surface of sandy substrata, up to 16 mm diameter; many colonies broken through predation or physical disturbance; irregular fragments regenerating to repair missing areas and continuing to grow. Convex upper surface of the colony consisting of rows of radially arranged zooids (Fig. 11 A – D). Autozooids rhomboidal, with frontal membrane underlain laterally and proximally by granular cryptocyst. At distal end of each zooid an interzooecial vibraculum with an ear-shaped chamber (Fig. 11 C, D) and a mandible in the form of a long curved seta. Large vicarious vibracula, with scimitar-shaped setae occur in ancestrular region at apex of colony or in areas of regeneration. Lower surface of the colony consisting of extrazooidal calcification, which forms concentric rings, divided into small square sectors, each with 1 - 6, mostly 2 - 4, round pores (Fig. 11 E, F). No ooecia. Embryos brooded in zooids."},{"description":"Cupuladria biporosa Canu & Bassler, 1923: 29, pl. 47, figs 1 – 2; Cook 1965: 203, pl. 1, figs 2 A, B, 3 A, B, 4 A, B, 5, 6 A, B, text-fig. 1, g-j; Winston 2005: 13, figs 30, 37. Cupuladria canariensis: Canu & Bassler 1928 a: 16 (part), text-fig. 2; Marcus & Marcus 1962: 285, pl. 1, figs 1 – 2; Lagaaij 1963 b: 225 (part), pl. 26, figs 4, 5. Cupuladria sp. 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Contributions to Canadian Biology and Fisheries New Series, 1: 143–201. https://www.marinespecies.org/aphia.php?p=sourcedetails&id=300150","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1010029","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","53":"Bryozoa","252":"Gymnolaemata","467":"Cheilostomatida","6020":"Schizoporellidae","1007532":"Schizoporella"},"class":"Gymnolaemata"},{"key":1010040,"nameKey":10154686,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"2d59e5db-57ad-41ff-97d6-11f5fb264527","nubKey":1010040,"parentKey":1007532,"parent":"Schizoporella","basionymKey":9812207,"basionym":"Eschara biaperta Michelin, 1848","kingdom":"Animalia","phylum":"Bryozoa","order":"Cheilostomatida","family":"Schizoporellidae","genus":"Schizoporella","species":"Schizoporella biaperta","kingdomKey":1,"phylumKey":53,"classKey":252,"orderKey":467,"familyKey":6020,"genusKey":1007532,"speciesKey":1010040,"scientificName":"Schizoporella biaperta (Michelin, 1848)","canonicalName":"Schizoporella biaperta","authorship":"(Michelin, 1848) ","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1010040","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","53":"Bryozoa","252":"Gymnolaemata","467":"Cheilostomatida","6020":"Schizoporellidae","1007532":"Schizoporella"},"class":"Gymnolaemata"},{"key":1010065,"nameKey":10154848,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1010065,"parentKey":1007532,"parent":"Schizoporella","basionymKey":6487220,"basionym":"Cellepora dunkeri Reuss, 1848","kingdom":"Animalia","phylum":"Bryozoa","order":"Cheilostomatida","family":"Schizoporellidae","genus":"Schizoporella","species":"Schizoporella dunkeri","kingdomKey":1,"phylumKey":53,"classKey":252,"orderKey":467,"familyKey":6020,"genusKey":1007532,"speciesKey":1010065,"scientificName":"Schizoporella dunkeri (Reuss, 1848)","canonicalName":"Schizoporella dunkeri","authorship":"(Reuss, 1848) ","publishedIn":"World List of Bryozoa. 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It is also very abundant in several areas including the northwest Iberian Peninsula (Reverter-Gil & Fernández-Pulpeiro 1999, 2001). In the Mediterranean, despite difficulties with synonymy (see Hayward & Ryland 1995, 1999), it seems to be abundant from shallow water to 60 m and it is very common in the Adriatic (Hayward & McKinney 2002). Our findings represent the first record of S. dunkeri for Madeira."},{"description":"The species is able to colonise stones and shells. It occurs in subtidal waters and has been recorded from 12-60 metres."},{"description":"Schizoporella dunkeri is distributed from the Mediterranean northwards to the Isles of Scilly. It is abundant throughout the Adriatic in shallow environments with a high detritus load."},{"description":"Schizoporella dunkeri is an encrusting bryozoan that forms extensive spreading sheets, composed of a single, or multiple layers of autozooids. Colonies are reddish to orange in colour. Autozooids are large, very broad and roughly rectangular. 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(1830). Ueber einige fossile Arten Cypris (Mueller, Lamk.) und Cythere (Mueller, Latreille, Desmarest). 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A. (1946). Contribution to the Study of Ostracoda with Special Reference to the Tertiary and Cretaceous Microfauna of the Caribbean Region. Amsterdam, DeBussy, 1–167. https://www.marinespecies.org/ostracoda/aphia.php?p=sourcedetails&id=173514","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1028911","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Marginal pores In Triebelina sp., six large marginal spines are visible along the postero-ventral margin (Fig. 9 A, B). They are hollow, but no setae are associated with them. A tiny pore is also visible at the base of each spine (Figs 5 A; 9 B), which houses the marginal setae (eyelash setae ofMaddocks 2013). The associated radial pore canals cannot be tracked through the entire thickness of the valve wall because of low resolution. The two adult specimens of Triebelina indopacifica are overall more worn, and marginal spines are less well preserved, although they have been illustrated on the type material (van den Bold 1946: fig. 7). Four broken marginal spines occur along the antero-ventral margin of the smaller LV, and minute structures along the posteroventral margin (Fig. 4). Lateral normal pores In all three Triebelina specimens, lateral NPC cross the entire thickness of the valve perpendicularly to the surface (Figs 3 H, I; 5 H, I). They are not uniformly distributed, as they are closely associated with macro-ornamentation and less abundant in uniquely micro-ornamented zones (Figs 3 J; 4 H; 5 J). On the scans and 3 D reconstructions, these macro-ornamentation related canals are rarely observed to cross the thickest portion of ornamentation features, while they are more abundant on the SEM images (Figs 3 A; 4 A; 5 A). These differential observations are, therefore, related to the resolution of the CT-scan, small pore canals being undetected. The following preliminary description consequently focuses on the largest pore canals clearly observable through scans. Two rows of pore canals border the ventral ridge of all specimens and the median bulge in the case of T. indopacifica (Figs 3 J; 4 H; 5 J). Dorsally, they are also distributed into two rows along the horizontal ridge as well as the anterior and posterior lateral vertical elements. In areas lacking macro-ornamentation, pores are mainly located at the anterior and posterior ends of the lateral surface, where the crests and ventral ridge interrupt, but this pattern is strongly biased by the resolution limits of the CT-scan. The smallest LV of T. indopacifica displays numerous pore canals through the thickest parts of the macro-ornamentation, mainly through the posterior portion of the ventral ridge, while they are less numerous near the AMS (Fig. 4 H, I). Overall, a certain polarity of the distribution of canals for all the three valves studied is visible, with denser distribution along the ventral margin compared to the dorsal area. The smaller LV of T. indopacifica displays a unique set of nine double pore systems aligned in two rows above the anterior termination of the ventral ridge (Fig. 4 H-J). At the time of writing, we are not aware of any previous work reporting such systems of double pore canals in ostracods. These canals are relatively narrow, and they cross the valve thickness while being very closely positioned, nearly touching each other. The possibility that they may correspond to artifacts is precluded, as their respective walls are clear and coherent in axial, coronal and sagittal views. One anastomosed pore canal is also seen at the posterior end of the ventral ridge of the same LV (Fig. 4 I, K). Such unique pore systems, their relation to setae and calcification, and taxonomic significance should be appropriately investigated in the future. Overall, the area of the ventral ridge in T. indopacifica here appears as the richest in terms of diversity of pore canal morphologies. The position of pores regarding reticulation is shown in Figs 3 J; 4 H; 5 J. The reticulation at the surface of all macroornamental areas (dorsal, ventral and median ridges) is blurred and it is impossible to determine the relationship of pores to reticulation. In other areas, pores in all three Triebelina specimens are intramural. Further analysis of juvenile specimens will be of importance to compare the ontogeny of pore systems with the ontogenetic history of calcification. The study of complete carapaces will clarify how pore canals are organized in the two valves of these highly asymmetrical ostracods, if the asymmetry of the valves also corresponds to an asymmetry of the pore systems."},{"description":"The present 3 D reconstructions illustrate the major morphological characters of T. indopacifica. The two largest valves have a wide calcified inner lamella and are adult (Figs 3 C; 4 C). In both specimens, the AMS are located within the cavity underlying the swelling of the ventro-lateral ridge but are weathered and cannot be satisfactorily observed. The RV (Fig. 5) is small, thin-walled, and has a narrow calcified inner lamella; it is an instar. The AMS pattern is plain, consisting of eight elongate scars organised in four parallel diagonal rows (Fig. 5 C). The species identification of this juvenile is uncertain."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","353":"Ostracoda","1438":"Podocopida","5190":"Bairdiidae","1028907":"Triebelina"},"class":"Ostracoda"},{"key":1028948,"nameKey":6586955,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1028948,"parentKey":1028942,"parent":"Macrocyprina","basionymKey":1029376,"basionym":"Paracypris pacifica LeRoy, 1943","kingdom":"Animalia","phylum":"Arthropoda","order":"Podocopida","family":"Macrocyprididae","genus":"Macrocyprina","species":"Macrocyprina pacifica","kingdomKey":1,"phylumKey":54,"classKey":353,"orderKey":1438,"familyKey":5207,"genusKey":1028942,"speciesKey":1028948,"scientificName":"Macrocyprina pacifica (Leroy, 1943) Swain, 1967","canonicalName":"Macrocyprina pacifica","authorship":"(Leroy, 1943) Swain, 1967","publishedIn":"Leroy, L. W. (1943). 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A. (1966). Miocene and Pliocene Ostracoda from northeastern Venezuela. Verh.Kon.Akad.Wet.Amsterdam, Afd.Natuurk.(1, 23: 5–43. https://www.marinespecies.org/ostracoda/aphia.php?p=sourcedetails&id=173393","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1028966","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","353":"Ostracoda","1438":"Podocopida","9272":"Pontocyprididae","1028965":"Argilloecia"},"class":"Ostracoda"},{"key":1028977,"nameKey":9098882,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"9ca92552-f23a-41a8-a140-01abaa31c931","nubKey":1028977,"parentKey":1028976,"parent":"Pontocypris","kingdom":"Animalia","phylum":"Arthropoda","order":"Podocopida","family":"Pontocyprididae","genus":"Pontocypris","species":"Pontocypris edwardsi","kingdomKey":1,"phylumKey":54,"classKey":353,"orderKey":1438,"familyKey":9272,"genusKey":1028976,"speciesKey":1028977,"scientificName":"Pontocypris edwardsi","canonicalName":"Pontocypris edwardsi","authorship":"","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1028977","extinct":true,"habitats":["MARINE"],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","353":"Ostracoda","1438":"Podocopida","9272":"Pontocyprididae","1028976":"Pontocypris"},"class":"Ostracoda"},{"key":1029376,"nameKey":8156631,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1029376,"parentKey":1028942,"parent":"Macrocyprina","acceptedKey":1028948,"accepted":"Macrocyprina pacifica (Leroy, 1943) Swain, 1967","kingdom":"Animalia","phylum":"Arthropoda","order":"Podocopida","family":"Macrocyprididae","genus":"Macrocyprina","species":"Macrocyprina pacifica","kingdomKey":1,"phylumKey":54,"classKey":353,"orderKey":1438,"familyKey":5207,"genusKey":1028942,"speciesKey":1028948,"scientificName":"Paracypris pacifica LeRoy, 1943","canonicalName":"Paracypris pacifica","authorship":"LeRoy, 1943","publishedIn":"Leroy, L. W. (1943). Pleistocene and Pliocene Ostracoda of the Coastal Region of Southern California. 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grimaldii","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1470,"familyKey":5602,"genusKey":1142263,"speciesKey":1142316,"scientificName":"Plectromerus grimaldii Nearns & Branham, 2005","canonicalName":"Plectromerus grimaldii","authorship":"Nearns & Branham, 2005","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1142316","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Type. Holotype, female, in the collection of the AMNH, No. DR 16 535. Included in a piece of Dominican amber (OligoMiocene) from the Dominican Republic. Amber yellowbrownish, moderately clear; cut and polished to a flat, oval shape, measuring 18.5 X 15 X 8 mm. Specimen is in good condition except damage to left antenna: antennomere 7 is incomplete, antennomeres 8 – 11 are missing."},{"description":"Etymology. We are pleased to name this species for Dr. David Grimaldi (AMNH) for his contributions to the study of insects in amber and for making the specimen of this species available for study."},{"description":"Discussion. Although gender cannot be determined conclusively, we believe the holotype of P. grimaldii to be female based on the evenly, broadly rounded fifth visible abdominal ventrite and the lack of an irregular patch of coarse punctures in front of each prosternal coxa (a male characteristic seen in many extant species of Plectromerus). From other congeners, P. grimaldii can be distinguished by the following combination of characters: the shape and punctation of pronotum (widest at middle, alveolatepunctate), the elytral punctation (dense, coarse), the glabrous pronotum and elytra, and the small, nonserrate metafemoral tooth (Fig. 2 c). Plectromerus punctatus (Fisher) and P. exis Zayas also have small metafemoral teeth which are not serrate, however these species can be distinguished by having the third antennomere longest (fifth longest in P. grimaldii) and different pronotal dimensions: in P. punctatus the pronotum is almost as wide as long, in P. exis the pronotum has a distinct tubercle in the center and the length is about 1.8 times the width (1.5 times as long as wide in P. grimaldii). Notes on Plectromerus tertiarius Vitali holotype: ventral habitus as in Fig. 1 b (dorsal habitus completely obscured), length approximately 7 mm (exact measurement not possible since abdomen is bent up through open elytra), included in a piece of Dominican amber (Lower Miocene) from the Dominican Republic. Amber yellowbrownish, partially obscured by numerous, small bubbles; cut and polished in a nearoval shape, measuring 42 X 22 X 15 mm. Specimen is damaged as follows: metathoracic legs are missing except coxae and trochanters; left antenna is damaged, missing part of antennomere 8, completely missing antennomeres 9 – 11. One important character in particular, the prosternal process between coxae, is not visible due to position of pro and mesothoracic legs. Elytral punctation can be inferred from ventral view due to open elytra which are semitranslucent. Vitali (2004) states that the holotype is a male, however, we see nothing to support this. In our opinion, the broadly rounded fifth abdominal segment is more indicative of a female Plectromerus (irregular patches of coarse punctures in front of each prosternal coxa are also not visible but the view is partially obscured). Vitali (2004) also states that the first abdominal ventrite is 3 times longer than other visible ventrites, however, our measurements show it to be about 2 times longer."},{"description":"Plectromerus grimaldii superficially resembles P. tertiarius in pronotal shape and elytral punctation. They differ, however, with respect to elytral apices (subtruncate in P. grimaldii, evenly rounded in P. tertiarius) and visible abdominal ventrite relationships (first ventrite as long as next 3 visible abdominal ventrites combined in P. grimaldii, first ventrite slightly longer than next 2 visible abdominal ventrites combined in P. tertiarius). In addition, significant differences can be seen in antennomere morphology. These differences exceed the variation in antennal morphology seen in extant species and across gender in Plectromerus. In P. grimaldii, the fifth antennomere is about 1.9 times longer than the tenth (about 1.6 times longer in P. tertiarius), fifth antennomere about 1.5 times longer than seventh (about 1.1 times longer in P. tertiarius). In P. tertiarius, the seventh antennomere is slightly longer than the sixth (subequal in P. grimaldii) and the eleventh antennomere is slightly longer than the tenth (subequal in P. grimaldii). In addition, antennomeres 5 – 10 are distinctly produced externally in P. tertiarius, whereas in P. grimaldii, antennomeres 6 – 10 are only moderately produced externally (Fig. 3 b – d)."},{"description":"Description. FEMALE. Length 7.1 mm, width 1.8 mm (measured across humeri). Habitus as in Figure 1 a. General form small, narrow, subcylindrical. Elytra with two indistinctly defined and very faint, transverse ferruginous fasciae on each elytron, one at basal third and another just behind middle. Head with front nearly flat, transverse, with a median, shallow line from between eyes to just beyond vertex, slightly concave between antennal tubercles, which are somewhat raised and widely separated. Much of head surface is obscured by an opaque film, exposed areas with surface opaque, alveolatepunctate. Eyes coarsely faceted, prominent, transverse, subreniform (Fig. 2 b). Antennae elevensegmented, slightly longer than body, impunctate; scape bowed, third antennomere subequal to scape, almost twice as long as fourth, fifth antennomere longest, about 2.6 times longer than fourth, antennomeres 6 11 becoming progressively shorter, sixth through eighth slightly longer than third, eleventh slightly longer than fourth, basal antennomeres subcylindrical, from fifth slightly flattened, apices of antennomeres 6 – 10 produced externally, antennomeres 7 – 9 slightly bowed (Fig. 3 b, 3 d). Antennomeres 2 – 6 ciliate beneath with coarse, moderately long, suberect, hairs. Pronotum subcylindrical, about 1.5 times as long as wide, widest at middle, slightly broader at apex than base, sides broadly inflated, arcuately constricted at basal third, and a slight inflation just before apex; basal margin slightly arcuate; disk convex. Surface opaque, alveolatepunctate. Scutellum small, rounded, almost as long as broad, impunctate. Elytra about 2.7 times as long as width at humeri, about 2.5 times as long as pronotal length, about 1.3 times broader basally than pronotum at widest (at middle); sides nearly parallel, slightly sinuate around middle, evenly rounded to apex which is subtruncate; epipleural margin moderately sinuate. Elytral disk slightly concave medially, subsuturally, creating a faint costa on each elytron. Surface shining; punctation dense, coarse, punctures becoming finer towards apex and sides, almost obsolete on apical third; glabrous. Underside with prosternum shining; narrowest area of prosternal process between coxae about 0.25 times as wide as coxal cavity, and about 0.5 times the width of apex of process which is cordate (emarginated at middle of apex). Mesosternum surface shining, sparsely and finely punctate (Fig. 2 b). Metasternum surface shining, sparsely and finely punctate, with a few suberect hairs; first visible abdominal ventrite longest, about 2.5 times longer than second, about as long as next three visible abdominal ventrites combined, fifth visible abdominal ventrite evenly, broadly rounded, slightly longer than fourth. Legs very finely punctate, with femora clavate, meso and metafemora arcuate, underside of each femoral club with a small triangular tooth with posterior edge smooth; metafemora about 1.2 times longer than metatibiae; tibiae slightly flattened, expanded apically, base of tibiae slightly arcuate (Fig. 2 c)."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1470":"Coleoptera","5602":"Cerambycidae","1142263":"Plectromerus"},"class":"Insecta"},{"key":1144004,"nameKey":8155264,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1144004,"parentKey":1153592,"parent":"Stictoleptura","acceptedKey":6267953,"accepted":"Stictoleptura antiqua (Vitali, 2005)","kingdom":"Animalia","phylum":"Arthropoda","order":"Coleoptera","family":"Cerambycidae","genus":"Stictoleptura","species":"Stictoleptura antiqua","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1470,"familyKey":5602,"genusKey":1153592,"speciesKey":6267953,"scientificName":"Paracorymbia antiqua Vitali, 2005","canonicalName":"Paracorymbia antiqua","authorship":"Vitali, 2005","publishedIn":"VITALI Francesco. Notes about the european fossil Lepturinae and the description of a new species (Coleoptera, Cerambycidae, Lepturinae). Lambillionea 105 (4) 1: 530-538, 7 figs. 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(1758): Systema naturae. Regnum Animale. 10 th ed. Lipsiae: W. Engelmann, URL: http://hdl.handle.net/10199/15420","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1315083","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Hab. Europe and North America."},{"description":"Hab. Throughout Europe. Madeira."},{"description":"Habitat in Europae terra."},{"description":"F. tota nigra nitida, tibiis cinerascentibus."},{"description":"Obs. Hanc F. nigram L. veram esse testantur definitio et locus (\" habitat sub terra \" 1. c), quod minime in F. fuliginosam convenit, ad quam refert, Cel. Dahlbom, nomen F. nigrae L. Quid vero revera sit F. obsoleta Linn. Fn. Sv. 1724, hoc explicare in mea potestate non est, nec quomodo ad F. nigrum nostram, ut in mscrpto suo vult Cel. Dahlbom, referatur, intelligo. In F. ebsoletam L. valent quoque, ex mea sententia, verba Latreillei: \" Linnne a decrit quelques insectes d'une maniere si legere, qu'on est oblige de se mettre l'esprit a la torture pour les reconnoitre. \" C. Flavae: colore operariarum saltem dominante flavo."},{"description":"[[ worker ]] Long. 1 1 / 4 - 1 1 / 2 lin. Caput nitidum sericeo pubescens, mandibulis, genis scapisque sordide testaceo-rufescentibus; palpi longiusculi prothoracem attingentes; clypeus fere absque vestigio cannulas cum area triangulan confluens, limite tantum obsoletissimo, Oculi subimmersi vel vix supra superficiem temporum prominuli (nec ut in F. glebaria prominuli), ocelli minutissimi vel vix ulli. Thorax sicut in praecedente, metanoto tarnen breviori et pilis nonnullis flavidis sparsis erectis tamquam etiam in capite, praeter pubescentiam adpressam sericeam. Pedes fusci cinereo-pubescentes tenuissimeque albo-pilosuli, articulationibus et tarsis ferrugineo-pallescentibus. Squama parva verticaliter subrectangularis, margine supero parum vel vix emarginato. Abdomen ovatum sparse flavido-setulosum, marginibus segmentorum summis obsolete membranaceo-cinerascentibus. [[ queen ]]. Long. 3 1 / 2 lin. tota dense cinerascenti-sericeo pubescens. Caput paribus fere ut in [[ worker ]]. Oculi tennuiter obsolete parce pilosuli; ocelli sat magni. Thorax capite latior, pilis ut hoc parvis cinerascentibus erectis parcius conspersus. Alae hyalinissimae albo conspicae tinctae, nervis et stigmate cinerascentibus, radice et tegula fuscis-, alae anticae 4 lin. longae. Pedes ut in diagnosi colorati, tibiis inprimis dense cinerascenti-flavido-pubescentibus. Squama valde compressa verticaliter subrectangularis etcet ut in diagnosi. Abdomen longitudine fere capitis thoracisque cum petiolo, oblongo-ovale, segmentornm marginibus summis tenuissime membranaceo-pallescentibus; pilis parvis solitis, prater pubescentiam densam sericea m aequaliter undique vestientem. [[ male ]]. Long. circ. 1 3 / 4 lin. Dens unicus mandibularum apice rufescens. Antennae nigrofuscas, articulo flagellorum primo sive pedicello breviter obconico, conspicue crassiori quam ceteri articuli. Oculi nudi. Linea frontalis profunde impressa. Occiput parum concaviusculum. Alae nervis et stigmate dilutius cinerascentibus quam in femina; anticae fere 2 1 / 2 lin, longae, area discoidali parva sub quadrata vel nulla. Squama parva compressa, ut in diagnosi. Abdomen longitudine thoracis supra visum ovatum nitidum nigrum, marginibus segmentorum summis tenuissime cinerascentibus; vaginarum genitalium binis paribus exterioribus fuscis brevioribus, pari medio albido longiori, omnibus linearibus, apicibus rotundatim obtusis, externis pilosis; valvula ventrali semiovali parva pilis parcis."},{"description":"Operaria: fusco-nigrescens cinerascenti-micans nitida, sparse flavido pilosula, mandibulis et antennarum scapis rufescentibus, tarsis testaceo-pallescentibus; ocellis minutissimis; squama parva subrectangulari supra parum vel vix emarginata. Femina: fusco-nigrescens dense undique cinereo-micans nitida, mandibulis antennarumque scapis obscure rufescentibus, pedum articulalionibus tarsisque ferrugineo-pallescentibus; alis albohyalinis nervis et stigmate pallide flavidis; squama verticaliter subrectangulari, supra angulis rotunda tis et medio subangulatim emarginato. Mas: fusco-niger parum cinereo-micans nitida, tarsis obscure pallescentibus; flagellorum articulo primo crassiusculo; squama parva transversim subrectangulari supra parum concaviuscula; vaginis genitalium linearibus, pari interno (vel medio) albido, longiori quam externo."},{"description":"Hab. per terras nostras usque in Lapponiam vulgaris, sub lapidibus, cortice arborum et in arena inprimis nidulans. Examinat apud nos a fine niensis Julii ad initium usque Septembris."},{"description":"FORMICA NIGRA HNS. Formica nigra, Linn HNS. Faun. Suec. 1723. Nyland. Adno. Mon. Form. Boreal. 920, 16. Formica fusca, Foerster HNS, Hym. Stud. Form. 33, 14."},{"description":"The Garden Ant. B. 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Figs 1 A, B, 2 A, B. Body length 5.5 mm. Head width not properly measurable in specimen, but head appears slightly elongated, with almost straight and subparallel sides, narrowly rounded occipital corners and straight occipital margin. Eyes of moderate size, elongate-oval, situated in front of midlength of sides of head. Ocelli barely visible due to nature of sculpture (see below). Frontal carinae absent; frontal lobes weakly developed, horizontal, antennal sockets mostly exposed and toruli not concealed by frontal lobes; antennal scrobes absent. Clypeus posteriorly rather widely inserted between frontal lobes, its median part flattened, delineated laterally by coarse longitudinal carinae, with three longitudinal costae each, terminated at the apices with sharp teeth; lateral parts of clypeus raised into a ridge in front of toruli; anterior clypeal margin with paired long setae, without medial seta, its median part almost straight. Mandibles rather long, with distinct masticatory margin, armed with two large sharp apical teeth followed by two blunt tuberculate denticles. Maxillary palps 4 - segmented, labial palps 3 - segmented. Antennae 12 - segmented, with distinct 3 - segmented club. Antennal scape gradually curved at base and with no trace of lobe or carina, relatively short, far not reaching occipital margin. First funicular segment ca. twice as long as wide, 2 nd – 8 th segments transverse, total length of 9 th – 11 th segments subequal to total length of preceding segments. Mesosoma short, high, and robust. Pronotum well developed, scutum slightly convex, scutellum flattened. Propodeal dorsum feebly convex, somewhat shorter than posterior surface, propodeal lobes slightly angulated but not pointed. Propodeal spines widened at the base, very long, ca. half of head length, curved down along their length, sharply pointed at apices, and subparallel (as seen from above). Petiole of moderate length and quite low, PL / PH 1.74, PL / HL 0.45; length of anterior peduncle subequal to length of node; anterior surface of node concave, its dorsum rounded, posterior surface somewhat convex, gradually sloping backwards; petiole ventrally with longitudinal lamella. Postpetiole subglobular, 1.35 times as wide as petiole. Legs of moderate length (HTL / ML 0.51, HFL / ML 0.67), femora and tibiae somewhat incrassated, pretarsal claws simple, arolia well developed. Meso- and metatibiae with well-developed simple spur. Body coarsely sculptured. Entire head (except for clypeus), pronotum, scutum, scutellum and propodeal dorsum with big and deep foveae, that touch each other; mesopleura and sides of propodeum with very coarse longitudinal costae; petiolar node and postpetiole with less coarse costulate sculpture; base of first gastral tergite near postpetiole with short longitudinal costulae; gastral tergites with scattered piligerous pits or small foveae. Mandibles with longitudinal rugae. Head and mesosoma with sparse, long erect and short suberect setae; petiolar node dorsum with two suberect setae; shorter suberect setae are visible only on ventral surface of postpetiole. All gastral sternites with quite numerous and long subdecumbent to suberect setae; first gastral tergite appears bare, pilosity on remaining tergites are hidden by whitish coating. Antennae and legs with abundant short subdecumbent pilosity. Measurements (in mm) and ratios: HL 1.14, SL 0.73, OL 0.22, OW 0.17, GL 0.16, MdL 0.60, ML 1.46, MH 0.94, PL 0.52, PH 0.30, PW 0.26, PPL 0.39, PPH 0.33, PPW 0.35, HTL 0.74, HFL 0.98, ESL 0.55, ESD 0.52; SL / HL 0.64, OL / HL 0.19, OL / OW 1.31, OL / GL 1.42, MdL / HL 0.52, PL / PH 1.74, PL / PW 2.00, PL / HL 0.45, PPL / PPH 1.20, PPW / PW 1.35, ESL / HL 0.48, ESD / ESL 0.95, ML / MH 1.56, HTL / ML 0.51, HFL / ML 0.67. Males unknown. The studied queen possesses all the diagnostic features of the Stiphromyrmex robustus workers, differing from them by the structure of the mesosoma and sculpture of the mesopleura and sides of the propodeum, which are longitudinally coarsely costulate (compare Fig. 1 and Fig. 3). It undoubtedly belongs to this genus and most likely to S. robustus. However, we do not exclude that it may belong to another, as yet undescribed species of this genus, but we have no arguments in favor of this assumption. Therefore, for now, we classify it as Stiphromyrmex robustus."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1457":"Hymenoptera","4342":"Formicidae","1318998":"Stiphromyrmex"},"class":"Insecta"},{"key":1319138,"nameKey":8195542,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1319138,"parentKey":1319137,"parent":"Paraneuretus","kingdom":"Animalia","phylum":"Arthropoda","order":"Hymenoptera","family":"Formicidae","genus":"Paraneuretus","species":"Paraneuretus tornquisti","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":4342,"genusKey":1319137,"speciesKey":1319138,"scientificName":"Paraneuretus tornquisti Wheeler, 1915","canonicalName":"Paraneuretus tornquisti","authorship":"Wheeler, 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The species, represented by the major worker, is identifiable as a member of the Ph. megacephala species group by (1) the presence of the conspicuous ventral convexity of the postpetiolar sternum (Fig. 19 A; e. g., Salata and Fisher 2020). It differs from Ph. megacephala (Fabricius, 1793), Ph. megatron Fischer & Fischer, 2013, and Ph. spinosa Forel, 1891 by (2) the well-developed inner hypostomal teeth (Fig. 20 B; e. g., Salata and Fisher 2022). Among the Pheidole megacephala group species more broadly (e. g., Fischer et al. 2012), it differs in having (3) facial rugosity that extends to the posterior margin of the occipital lobes (Fig. 20 A, note: among type specimens of the group imaged on AntWeb, this condition also occurring in Ph. megacephala impressifrons Wasmann, 1905, which has a more angular bulge of the postpetiolar sternum). See the description below for further conditions."},{"description":"Figs 19, 20, 21: Fig. A 5"},{"description":"Description. Measurements (in mm; abbreviations follow Salata and Fisher 2022): EL = 0.152; HL = 1.290; HW = 1.230; MTL = 0.673; PNW = 0.618; PPW = 0.324; PSL = 0.202; PTW = 0.172; SL = 0.714; WL = 1.110. Indices (also following Salata and Fisher 2022): CI = 95.3; MTI = 54.7; SI = 58.0; PNI = 50.2; PPI = 26.3; PSLI = 16.4. Note: Measurements taken from cross-sectional projections in DragonFly using the reregistration and ruler tools. Head. In full-face view (Figs 19 B, 20 A), the head is subcordate, with the lateral margins widest somewhat beyond head midlength and with the posterior portions of the lateral margins converging posterad to the occipital lobes. In lateral view (Fig. 19 A), the head is subovate. The antennal scrobes are indistinct. The occipital lobes are rugose, with shagreened interspaces. The inner hypostomal teeth are well-developed; they are distant from the outer teeth, which are also well-developed (Fig. 20 B). The median hypostomal tooth is indistinct. Mesosoma. The humeral tubercle of the pronotum is weakly developed. The mesonotal bulge is distinct but not pronounced. The metanotum is only weakly indicated by a slight angularity of the promesonotal profile in lateral view. The propodeal spines are moderately long, with a wide base and acute tip (Fig. 19 A). Metasoma. The bulge of the postpetiolar sternum is rounded anteriorly. The first gastral tergum (ATIV) appears to be shagreened at its base (Fig. 20 D). Setation. Length and stature of setation uncertain, although density measurable in the scans based on the distinct occurrence of the setiferous punctation. Coloration. Not clearly visible; appears brownish / reddish."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1457":"Hymenoptera","4342":"Formicidae","1321654":"Pheidole"},"class":"Insecta"},{"key":1322533,"nameKey":8543717,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1322533,"parentKey":1321654,"parent":"Pheidole","kingdom":"Animalia","phylum":"Arthropoda","order":"Hymenoptera","family":"Formicidae","genus":"Pheidole","species":"Pheidole tethepa","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":4342,"genusKey":1321654,"speciesKey":1322533,"scientificName":"Pheidole tethepa Wilson, 1985","canonicalName":"Pheidole tethepa","authorship":"Wilson, 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conservatus","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":4342,"genusKey":1326115,"speciesKey":1326171,"scientificName":"Cerapachys conservatus Viehmeyer, 1913","canonicalName":"Cerapachys conservatus","authorship":"Viehmeyer, 1913","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1326171","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1457":"Hymenoptera","4342":"Formicidae","1326115":"Cerapachys"},"class":"Insecta"},{"key":1326284,"nameKey":4681371,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1326284,"parentKey":1326282,"parent":"Gesomyrmex","kingdom":"Animalia","phylum":"Arthropoda","order":"Hymenoptera","family":"Formicidae","genus":"Gesomyrmex","species":"Gesomyrmex hoernesi","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":4342,"genusKey":1326282,"speciesKey":1326284,"scientificName":"Gesomyrmex hoernesi Mayr, 1868","canonicalName":"Gesomyrmex hoernesi","authorship":"Mayr, 1868","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1326284","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Types. Mayr (1868) described G. hoernesi from 18 workers and a male. Three workers are preserved in NMW (1984 / 34 / 141, 1984 / 34 / 142 and 1984 / 34 / 143). A lectotype was not designated. The holotype of Dimorphomyrmex theryi is present at the collection of BMNH (In. 29052). The types of Gesomyrmex annectens and Dimorhomyrmex mayri are lost. Studied material. Baltic amber (workers). BMNH. In. 17787 with label \" Samland \"; BMNH. In. 29052 with label \" Coll. A. Thery, holotype of Dimorphomyrmex theryi Emery \"; GMUG. BST. 03830 (K 2614) determined by Wheeler (1929) as G. hoernesi; GMUG. BST. 03840 (K 2626); GMUG. BST. 03977 (228, K 4302); GMUG. BST. 03993 (K 4048); GMUG. BST. 04076 (K 6406); GMUG. BST. 04088 (a 181, K 4311); GMUG. BST. 04097 (x 10); GMUG. BST. 04151 (K 4442) determined by Wheeler as D. theryi; GMUG. BST. 04152 (K 4252) determined by Wheeler as D. theryi, pictured on fig. 50 in Wheeler (1915); GMUG. BST. 04157 (K 779) determined by Wheeler as D. theryi; GMUG. BST. 04200 (a 124) determined by Wheeler as G hoernesi; GMUG. BST. 04206 (a 119) determined by Wheeler as G hoernesi; GMUG. BST. 04207 (K 889) determined by Wheeler as G. hoernesi; GMUG. BST. 04208 (a 111) determined by Wheeler as G hoernesi; GMUG. BST. 04213 (K 4466) determined by Wheeler as G hoernesi; GMUG. BST. 04214 (a 95) determined by Wheeler as G hoernesi; GMUG. BST. 04438 (K 6421) determined by Wheeler as D. theryi; GMUG. BST. 04456 (a 204) determined by Wheeler as G hoernesi; GMUG. BST. 04457 (a 171) determined by Wheeler as G. hoernesi; MZPW. 412; MZPW. 2940; MZPW. 5864; MZPW. 5890; MZPW. 8126; MZPW. 9561; MZPW. 10353; MZPW. 15933; MZPW. 15980; MZPW. 19144; MZPW. 19377; MZPW. 19959; PIN. 964 / 484; PIN. 964 / 485; PIN 964 / 486; CGC. 1632; CGC. 2652; CGC. 3338. Bitterfeld amber. PMHU. 7 / 201 (worker); HM. 7 / 229 ([[ queen ]]); PMHU. 10 / 225 (worker); PMHU. 11 / 221 (worker); PMHU. 13 / 206 (worker); PMHU. 13 / 210 (worker); PMHU. 13 / 221 (worker); PMHU. 15 / 205 (worker); PMHU. 16 / 227; MKC. F- 010 [[ male ]]); MKC. F- 129 (worker); MKC. F- 130 (worker). Rovno amber (workers). SIZK. K- 419; SIZK. K- 5695. Scandinavian amber (workers). ZMUC. 187 with label \" G. Henningsen, 01.07.1966 \"; ZMUC. 188 with label \" G. V. Henningsen, 01.02.1969 \"; ZMUC. 189 with label \" G. V. Henningsen, 03.05.1960 \"; ZMUC. 190 with label \" A. K. Andersen, 28.03.1968 \"."},{"description":"Distribution and horizon. Baltic, Bitterfeld, Rovno and Scandinavian ambers. Late Eocene."},{"description":"Comments. There is a problem with the association of different castes of fossil ant species as findings of males, females and workers of the same species in the same piece of amber are extremely rare. However we are confident that workers, male and female of Gesomyrmex described above belong to the same species. Up to the present 231 workers of Gesomyrmex from Baltic and similar ambers are studied: 18 by Mayr (1868), 7 by Andre (1895), one by Emery (1905), 172 by Wheeler (1915) and 33 (excluding species determined by Wheeler) by one of us (GMD). All these specimens belong to the same species G hoernesi. So there is a high possibility that the two studied males (one by Mayr, and one by us) and above described gyne belong to the same species. The gyne has small eyes relative to those of the workers. This is unusual for ants, but Gesomyrmex is evidently an exception to the rule. The only extant species whose workers and gynes are known is G. luzonensis (Wheeler, 1916, 1930). Its gynes have comparatively smaller eyes than workers. Theobald (1937, p. 212) described Gesomyrmex hoernesi from Oligocene deposits of France (Kleinkembs, Haut-Rhin). Judged by the description and figures the described insects are really ants but they do not belong to Gesomyrmex. The male (Theobald 1937, pl. XIV, fig. 24) has a low nodiform petiole and filiform (not geniculate) antenna with a very short scape, and evidently belongs to Dolichoderinae. One of two specimens described as gynes (Theobald 1937, pl. XIV, fig. 25) has a rounded head and triangulate petiole without a scale. The form of the head of the other « gyne » (Theobald 1937, pl. XIV, fig. 26) is similar to Formica or Camponotus, but not to Gesomyrmex. This specimen also has a small and narrow petiole without a scale. Specimens described as workers (Theobald 1937, pl. XIV, figs. 27, 28) judged by the construction of mesosoma really are poorly preserved gynes without wings. They also have no characters of Gesomyrmex."},{"description":"Description. Minor worker. BL 2.5 - 5 mm. Body slender. Head longer than broad, subtrapezoid, narrower in the front than in the back, with convex sides, rounded posterior corners and feebly concave posterior border. Eyes very large, convex, reniform; long axes of eyes distinctive convergent anteriorly. Maximum eye diameter 1.8 - 2.0 times less than head length and 1.05 - 1.8 times less than head width. Cheeks very short. Ocelli small but distinct. Frontal area and frontal groove obsolete. Clypeus as long as broad or a little longer than broad. Anterior clypeal margin projecting as a long, rather narrowly rounded lobe over the bases of the mandibles. Antenna short, 8 - jointed. Scape does not reach the posterior margin of the eye. Funiculus incrassate, but without distinct club. First funicular joint more than twice as long than thick. The following joints longer than thick. Mandibles very long, decussating when closed, with 9 long, acute teeth, some of which are slightly shorter than the others. External margin of mandible concave in middle part. Maxillary palps do not reach the occipital foramen. Mesosoma long, narrower than the head, constricted in the mesopropodeal region. Pronotum gradually convex, longer than broad. Mesonotum rather flat above, the latter compressed laterally, longer than broad, narrower behind than in front and impressed, but without a suture that divides it from the metanotum. In lateral view propodeal dorsum and declivity are subequal and form a rounded obtuse angle; propodeal dorsum feebly convex, propodeal declivity slightly concave. Legs rather slender. Middle and hind tibiae each with one short simple spur. Petiole as long as broad, with low and thick scale, a little more than twice as broad as long, rounded above, with convex anterior and posterior surfaces. Gaster elongate elliptical, with a circlet of coarse anal cilia. Body rather shiny and lightly shagreened. Mandibles, clypeus and frons usually delicately longitudinally striated. Erect hairs absent on all parts of the body except the top of the gaster. Decumbent pubescence absent. Major worker (soldier). BL 5 - 8 mm. Head a little longer than broad, subrectangular, a little narrower in the front than in the back, with straight sides, rounded posterior corners and straight or feebly concave posterior border. Eyes large and convex, elongate elliptical or slightly reniform, situated near the middle of the sides, slightly more approximated in the front than in the back. Maximum eye diameter 2 - 3 times less than head length and 1.8 - 3.5 times less than head width. Ocelli usually lacking. Frontal carinae short but prominent, extending to the middle of the anterior orbits. Frontal area small and indistinct. Clypeus broader than long, its posterior border extending back medially between the frontal carinae. Anterior clypeal border projecting in the form of a short broad rounded lobe. Antenna short 8 - jointed. Scape not reaching to the posterior orbits. Funiculus incrassated but without distinct clave; joints 1 and 2 of the funiculi subequal, about twice as long as thick, joints 3 and 4 about 1.5 times as long as thick, 5 and 6 scarcely longer than thick, the terminal joint a little longer than thick. Mandibles stout, not decussating when closed, with convex external margin and 8 to 9 - toothed masticatory margin. The teeth are stout; the apical tooth is longest; shorter teeth are alternating with the longer ones towards the base. Maxillary palps short, do not reach the occipital foramen. Mesosoma long, narrower than the head, constricted in the mesopropodeal region. Pronotum gradually convex, about as long as broad. Mesonotum longer than broad and broader in front than behind; laterally its surface is straight and gradually sloping backward, its sides compressed. Metanotum distinct, short and transverse, divided from mesonotum and propodeum by sutures or impressions. In lateral view propodeal dorsum and declivity are subequal and form a rounded obtuse angle; propodeal dorsum feebly convex, propodeal declivity slightly concave. Legs long and rather stout, middle and hind tibiae with short simple spurs. Petiole short, with a low and thick scale. Gaster elongate oval. Body rather shiny and lightly shagreened. Mandibles, clypeus and frons sharply and densely longitudinally striated. Erect hairs absent on all parts of the body except the top of the gaster. Decumbent pubescence absent. Gyne (first description). BL 10.5 mm. Head subrectangular, about 1.3 times longer than wide, with rounded posterolateral corners and feebly concave posterior margin. Eyes comparatively smaller than in workers: maximum eye diameter less than 4 times head length. Ocelli large, distance between central and lateral ocelli equals roughly their diameters. Anterior clypeal margin and dentition of mandibles not visible. Antenna 10 - jointed. Scape does not reach the posterior margin. Funiculus incrassate, but without distinct club. First funicular joint longer than 2 and 3 together and 2.5 times longer than thick. Following joints except terminal about as long as thick. Scutum and scutellum wider than long. Gaster, petiole and propodeum covered by white film and not visible. Head and mesosoma with numerous short erect hairs. Legs without erect and semi-erect hairs. Male. BL 7 - 8 mm. Head with enormous large convex eyes, which occupy most of the head. Ocelli large and convex. Frontal carinae and lobes absent. Frontal furrow distinct. Anterior clypeal margin projecting as a rounded lobe. Antenna geniculate, small. Mayr (1868) wrote that it is 11 - jointed but the studied specimen has only one antenna with 8 joints; perhaps the terminal joints were lost. Mandibles short, not opposable, with an acute apical tooth and without teeth on the masticatory margin. Maxillary palps long, reach the occipital foramen. Pronotum transverse. Scutum with flat dorsal surface strongly rounded anteriorly. Scutellum convex. Propodeal dorsum and declivity form rounded obtuse angle in lateral view; propodeal dorsum much shorter than declivity. Legs rather long and thin. Petiole with rounded scale, higher than long. Gaster cylindrical. Genitalia not retractile. Body shiny with very delicate shagreened sculpture. Short erect hairs present on mandibles, clypeus, near ocelli, on the last sternites of the gaster, and genital parts. Decumbent pubescence absent. Forewing with closed cells 1 + 2 r, 3 r and mcu. Cell mcu rhomboidal. Veins RS and M partly united and form short section RS + M. Measurements (in mm). Minor workers. CGC. 1632: BL 5.0, AL 1.7, HL 1.2, MdL 0.64, ED 0.61, SL 0.58; CGC. 3338: BL 2.3, AL 0.73, HL 0.60, MdL 0.32, ED 0.31, SL 0.30; GMUG. BST. 03840: BL 4.7, AL 1.5, HL 1.0, ED 0.52, PtL 0.35. PtH 0.41; GMUG. BST. 04206: BL 4.4, AL 1.5, HL 1.0; GMUG. BST. 04207: BL 3.9, AL 1.3, HL 0.95, HW 0.68, MdL 0.50, ED 0.50, SL 0.52; GMUG. BST. 04208: BL 4.1, AL 1.35, HL 1.05, HW 0.75, MdL 0.50, ED 0.58, SL 0.48; GMUG. BST. 04213: BL 5.0, AL 1.5; GMUG. BST. 04214: BL 4.35, AL 1.4, HL 1.1, ED 0.58; GMUG. BST. 04456: BL 4.8, AL 1.55, HL 1.1, HW 0.8, ED 0.58, SL 0.55; PIN. 964 / 484: HL 0.99, HW 0.56, MdL 0.53, ED 0.53, SL 0.51; PIN. 964 / 485: BL 5.5, AL 1.75, HL 1.2, HW 1.1, ED 0.61, SL 0.64, PtL 0.35, PtW 0.45, PtH 0.48; PIN. 964 / 486: BL 5.5, AL 1.75. Major workers. GMUG. BST. 04151: BL 5.7; GMUG. BST. 04152: BL 5.8, AL 1.7, HL 1.4, HW 1.3, MdL 0.57, ED 0.47, SL 0.53; GMUG. BST. 04157: BL 5.3, AL 1.9; PMHU. 13 / 221: BL 5.3; BMNH. In. 29052: BL 5.5, AL 1.75, HL 1.25, HW 1.25, ED 0.53, SL 0.50; SIZK. K- 419: AL ~ 1.8, HL 1.2, HW 1.1, ED 0.62, SL 0.58. Gyne. PMHU. 7 / 229: BL 10.5, AL ~ 3, HL 2.2, HW ~ 1.6, ED 0.46, SL 0.92, FWL 6.5. Male. MKC. F- 010: BL about 7, AL 3.0, HL 1.3, MdL 0.36, ED 1.01, SL 0.34, FWL 6.8."},{"description":"(Fig. 2 and Fig. 6 A-E)"},{"description":"Studied material. Baltic amber (workers). BMNH. In. 17787 with label “ Samland ”; BMNH. In. 29052 with label “ Coll. A. Thery, holotype of Dimorphomyrmex theryi Emery ”; GMUG. BST. 03830 (K 2614) determined by Wheeler (1929) as G. hoernesi; GMUG. BST. 03840 (K 2626); GMUG. BST. 03977 (α 228, K 4302); GMUG. BST. 03993 (K 4048); GMUG. BST. 04076 (K 6406); GMUG. BST. 04088 (α 181, K 4311); GMUG. BST. 0 4097 (x 10); GMUG. BST. 0 4151 (K 4442) determined by Wheeler as D. theryi; GMUG. BST. 04152 (K 4252) determined by Wheeler as D. theryi, pictured on fig. 50 in Wheeler (1915); GMUG. BST. 04157 (K 779) determined by Wheeler as D. theryi; GMUG. BST. 04200 (α 124) determined by Wheeler as G. hoernesi; GMUG. BST. 04206 (α 119) determined by Wheeler as G. hoernesi; GMUG. BST. 04207 (K 889) determined by Wheeler as G. hoernesi; GMUG. BST. 04208 (α 111) determined by Wheeler as G. hoernesi; GMUG. BST. 04213 (K 4466) determined by Wheeler as G. hoernesi; GMUG. BST. 04214 (α 95) determined by Wheeler as G. hoernesi; GMUG. BST. 04438 (K 6421) determined by Wheeler as D. theryi; GMUG. BST. 04456 (α 204) determined by Wheeler as G. hoernesi; GMUG. BST. 04457 (α 171) determined by Wheeler as G. hoernesi; MZPW. 412; MZPW. 2940; MZPW. 5864; MZPW. 5890; MZPW. 8126; MZPW. 9561; MZPW. 10353; MZPW. 15933; MZPW. 15980; MZPW. 19144; MZPW. 19377; MZPW. 19959; PIN. 964 / 484; PIN. 964 / 485; PIN 964 / 486; CGC. 1632; CGC. 2652; CGC. 3338. Bitterfeld amber. PMHU. 7 / 201 (worker); HM. 7 / 229 (Ƥ); PMHU. 10 / 225 (worker); PMHU. 11 / 221 (worker); PMHU. 13 / 206 (worker); PMHU. 13 / 210 (worker); PMHU. 13 / 221 (worker); PMHU. 15 / 205 (worker); PMHU. 16 / 227; MKC. F- 010 (3); MKC. F- 129 (worker); MKC. F- 130 (worker). Rovno amber (workers). SIZK. K- 419; SIZK. K- 5695. Scandinavian amber (workers). ZMUC. 187 with label “ G. Henningsen, 01.07.1966 ”; ZMUC. 188 with label “ G. V. Henningsen, 01.02.1969 ”; ZMUC. 189 with label “ G. V. Henningsen, 03.05.1960 ”; ZMUC. 190 with label “ A. K. Andersen, 28.03.1968 ”."},{"description":"Distribution and horizon. Baltic, Bitterfeld, Rovno and Scandinavian ambers. Late Eocene."},{"description":"Comments. There is a problem with the association of different castes of fossil ant species as findings of males, females and workers of the same species in the same piece of amber are extremely rare. However we are confident that workers, male and female of Gesomyrmex described above belong to the same species. Up to the present 231 workers of Gesomyrmex from Baltic and similar ambers are studied: 18 by Mayr (1868), 7 by André (1895), one by Emery (1905), 172 by Wheeler (1915) and 33 (excluding species determined by Wheeler) by one of us (GMD). All these specimens belong to the same species G. hoernesi. So there is a high possibility that the two studied males (one by Mayr, and one by us) and above described gyne belong to the same species. The gyne has small eyes relative to those of the workers. This is unusual for ants, but Gesomyrmex is evidently an exception to the rule. The only extant species whose workers and gynes are known is G. luzonensis (Wheeler, 1916, 1930). Its gynes have comparatively smaller eyes than workers. Théobald (1937, p. 212) described Gesomyrmex hoernesi from Oligocene deposits of France (Kleinkembs, Haut-Rhin). Judged by the description and figures the described insects are really ants but they do not belong to Gesomyrmex. The male (Théobald 1937, pl. XIV, fig. 24) has a low nodiform petiole and filiform (not geniculate) antenna with a very short scape, and evidently belongs to Dolichoderinae. One of two specimens described as gynes (Théobald 1937, pl. XIV, fig. 25) has a rounded head and triangulate petiole without a scale. The form of the head of the other « gyne » (Théobald 1937, pl. XIV, fig. 26) is similar to Formica or Camponotus, but not to Gesomyrmex. This specimen also has a small and narrow petiole without a scale. Specimens described as workers (Théobald 1937, pl. XIV, figs. 27, 28) judged by the construction of mesosoma really are poorly preserved gynes without wings. They also have no characters of Gesomyrmex."},{"description":"Description. Minor worker. BL 2.5 – 5 mm. Body slender. Head longer than broad, subtrapezoid, narrower in the front than in the back, with convex sides, rounded posterior corners and feebly concave posterior border. Eyes very large, convex, reniform; long axes of eyes distinctive convergent anteriorly. Maximum eye diameter 1.8 – 2.0 times less than head length and 1.05 – 1.8 times less than head width. Cheeks very short. Ocelli small but distinct. Frontal area and frontal groove obsolete. Clypeus as long as broad or a little longer than broad. Anterior clypeal margin projecting as a long, rather narrowly rounded lobe over the bases of the mandibles. Antenna short, 8 - jointed. Scape does not reach the posterior margin of the eye. Funiculus incrassate, but without distinct club. First funicular joint more than twice as long than thick. The following joints longer than thick. Mandibles very long, decussating when closed, with 9 long, acute teeth, some of which are slightly shorter than the others. External margin of mandible concave in middle part. Maxillary palps do not reach the occipital foramen. Mesosoma long, narrower than the head, constricted in the mesopropodeal region. Pronotum gradually convex, longer than broad. Mesonotum rather flat above, the latter compressed laterally, longer than broad, narrower behind than in front and impressed, but without a suture that divides it from the metanotum. In lateral view propodeal dorsum and declivity are subequal and form a rounded obtuse angle; propodeal dorsum feebly convex, propodeal declivity slightly concave. Legs rather slender. Middle and hind tibiae each with one short simple spur. Petiole as long as broad, with low and thick scale, a little more than twice as broad as long, rounded above, with convex anterior and posterior surfaces. Gaster elongate elliptical, with a circlet of coarse anal cilia. Body rather shiny and lightly shagreened. Mandibles, clypeus and frons usually delicately longitudinally striated. Erect hairs absent on all parts of the body except the top of the gaster. Decumbent pubescence absent. Major worker (soldier). BL 5 – 8 mm. Head a little longer than broad, subrectangular, a little narrower in the front than in the back, with straight sides, rounded posterior corners and straight or feebly concave posterior border. Eyes large and convex, elongate elliptical or slightly reniform, situated near the middle of the sides, slightly more approximated in the front than in the back. Maximum eye diameter 2 – 3 times less than head length and 1.8 – 3.5 times less than head width. Ocelli usually lacking. Frontal carinae short but prominent, extending to the middle of the anterior orbits. Frontal area small and indistinct. Clypeus broader than long, its posterior border extending back medially between the frontal carinae. Anterior clypeal border projecting in the form of a short broad rounded lobe. Antenna short 8 - jointed. Scape not reaching to the posterior orbits. Funiculus incrassated but without distinct clave; joints 1 and 2 of the funiculi subequal, about twice as long as thick, joints 3 and 4 about 1.5 times as long as thick, 5 and 6 scarcely longer than thick, the terminal joint a little longer than thick. Mandibles stout, not decussating when closed, with convex external margin and 8 to 9 - toothed masticatory margin. The teeth are stout; the apical tooth is longest; shorter teeth are alternating with the longer ones towards the base. Maxillary palps short, do not reach the occipital foramen. Mesosoma long, narrower than the head, constricted in the mesopropodeal region. Pronotum gradually convex, about as long as broad. Mesonotum longer than broad and broader in front than behind; laterally its surface is straight and gradually sloping backward, its sides compressed. Metanotum distinct, short and transverse, divided from mesonotum and propodeum by sutures or impressions. In lateral view propodeal dorsum and declivity are subequal and form a rounded obtuse angle; propodeal dorsum feebly convex, propodeal declivity slightly concave. Legs long and rather stout, middle and hind tibiae with short simple spurs. Petiole short, with a low and thick scale. Gaster elongate oval. Body rather shiny and lightly shagreened. Mandibles, clypeus and frons sharply and densely longitudinally striated. Erect hairs absent on all parts of the body except the top of the gaster. Decumbent pubescence absent. Gyne (first description). BL 10.5 mm. Head subrectangular, about 1.3 times longer than wide, with rounded posterolateral corners and feebly concave posterior margin. Eyes comparatively smaller than in workers: maximum eye diameter less than 4 times head length. Ocelli large, distance between central and lateral ocelli equals roughly their diameters. Anterior clypeal margin and dentition of mandibles not visible. Antenna 10 - jointed. Scape does not reach the posterior margin. Funiculus incrassate, but without distinct club. First funicular joint longer than 2 and 3 together and 2.5 times longer than thick. Following joints except terminal about as long as thick. Scutum and scutellum wider than long. Gaster, petiole and propodeum covered by white film and not visible. Head and mesosoma with numerous short erect hairs. Legs without erect and semi-erect hairs. Male. BL 7 – 8 mm. Head with enormous large convex eyes, which occupy most of the head. Ocelli large and convex. Frontal carinae and lobes absent. Frontal furrow distinct. Anterior clypeal margin projecting as a rounded lobe. Antenna geniculate, small. Mayr (1868) wrote that it is 11 - jointed but the studied specimen has only one antenna with 8 joints; perhaps the terminal joints were lost. Mandibles short, not opposable, with an acute apical tooth and without teeth on the masticatory margin. Maxillary palps long, reach the occipital foramen. Pronotum transverse. Scutum with flat dorsal surface strongly rounded anteriorly. Scutellum convex. Propodeal dorsum and declivity form rounded obtuse angle in lateral view; propodeal dorsum much shorter than declivity. Legs rather long and thin. Petiole with rounded scale, higher than long. Gaster cylindrical. Genitalia not retractile. Body shiny with very delicate shagreened sculpture. Short erect hairs present on mandibles, clypeus, near ocelli, on the last sternites of the gaster, and genital parts. Decumbent pubescence absent. Forewing with closed cells 1 + 2 r, 3 r and mcu. Cell mcu rhomboidal. Veins RS and M partly united and form short section RS + M. Measurements (in mm). Minor workers. CGC. 1632: BL 5.0, AL 1.7, HL 1.2, MdL 0.64, ED 0.61, SL 0.58; CGC. 3338: BL 2.3, AL 0.73, HL 0.60, MdL 0.32, ED 0.31, SL 0.30; GMUG. BST. 03840: BL 4.7, AL 1.5, HL 1.0, ED 0.52, PtL 0.35. PtH 0.41; GMUG. BST. 04206: BL 4.4, AL 1.5, HL 1.0; GMUG. BST. 04207: BL 3.9, AL 1.3, HL 0.95, HW 0.68, MdL 0.50, ED 0.50, SL 0.52; GMUG. BST. 04208: BL 4.1, AL 1.35, HL 1.05, HW 0.75, MdL 0.50, ED 0.58, SL 0.48; GMUG. BST. 04213: BL 5.0, AL 1.5; GMUG. BST. 04214: BL 4.35, AL 1.4, HL 1.1, ED 0.58; GMUG. BST. 04456: BL 4.8, AL 1.55, HL 1.1, HW 0.8, ED 0.58, SL 0.55; PIN. 964 / 484: HL 0.99, HW 0.56, MdL 0.53, ED 0.53, SL 0.51; PIN. 964 / 485: BL 5.5, AL 1.75, HL 1.2, HW 1.1, ED 0.61, SL 0.64, PtL 0.35, PtW 0.45, PtH 0.48; PIN. 964 / 486: BL 5.5, AL 1.75. Major workers. GMUG. BST. 04151: BL 5.7; GMUG. BST. 04152: BL 5.8, AL 1.7, HL 1.4, HW 1.3, MdL 0.57, ED 0.47, SL 0.53; GMUG. BST. 04157: BL 5.3, AL 1.9; PMHU. 13 / 221: BL 5.3; BMNH. In. 29052: BL 5.5, AL 1.75, HL 1.25, HW 1.25, ED 0.53, SL 0.50; SIZK. K- 419: AL ~ 1.8, HL 1.2, HW 1.1, ED 0.62, SL 0.58. Gyne. PMHU. 7 / 229: BL 10.5, AL ~ 3, HL 2.2, HW ~ 1.6, ED 0.46, SL 0.92, FWL 6.5. Male. MKC. 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(1910).","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1359422","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1457":"Hymenoptera","9438":"Agaonidae","1359417":"Tetrapus"},"class":"Insecta"},{"key":1360322,"nameKey":6288675,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1360322,"parentKey":1360196,"parent":"Leucospis","kingdom":"Animalia","phylum":"Arthropoda","order":"Hymenoptera","family":"Leucospidae","genus":"Leucospis","species":"Leucospis glaesaria","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":9443,"genusKey":1360196,"speciesKey":1360322,"scientificName":"Leucospis glaesaria Engel, 2002","canonicalName":"Leucospis glaesaria","authorship":"Engel, 2002","publishedIn":"Engel, M.S. The first leucospid wasp from the fossil record (Hymenoptera: Leucospidae). JOURBOOK: Journal of Natural History VOLUME: 36(4) PAGES: 435-442. (2002).","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1360322","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Holotype. Female (figure 1), early Miocene amber, Dominican Republic, AMNH, DR- 10 - 186 9 (specific mine unknown). Labelled`Holotype, Leucospis glaesaria Engel. ’ Deposited in the amber fossil collection, Department of Entomology, American Museum of Natural History, New York, USA."},{"description":"Etymology. The specific epithet is taken from the Latin; glaesarius, meaning`of amber’."},{"description":"Diagnosis. The new species is distinctive for the combination of sparse pubescence, metallic cupreous integument on the mesosoma and metasoma, absence of a semicircular depression separating the lower tooth of the mandible (figure 3), a strong premarginal carina on the pronotum, carinate borders on the dorsellum, contiguous punctures on the mesosoma and metasoma (figure 2), a truncate metatibial apex (figure 5), long outer metatibial spur (figure 5), a long metabarsitarsus (figure 5), absence of a medial carina on the propodeum, a laterally conspicuous occipital carina, eight teeth on the ventral margin of the metafemur (figure 5), and a relatively short ovipositor (figure 4). It is most similar to L. speifera but differs by the absence of distinct interspaces between punctures, the cupreous body coloration, a depression along the posterior margin of the scutellum, absence of a medial carina on the propodeum and the sparse pubescence."},{"description":"Description. Female: Total body length (exclusive of antennae) 11.6 mm; forewing length 11.3 mm. Dorsal head width 3 mm, length 0.9 mm; frontal head width 3 mm, length 2.2 mm. Width of scrobes 0.6 mm. Clypeal length 0.5 mm, width 0.5 mm. Malar length 0.4 mm. Compound eye length 1.5 mm, width 1.2 mm; upper interorbital distance 1.5 mm; lower interorbital distance 1.4 mm. Interocellar distance (POL of BoucÏek) 0.4 mm; ocellocular distance (OOL of BoucÏek) 0.4 mm. Scape length 0.7 mm, width 0.2 mm; pedicel length 0.2 mm; flagellum length 4 mm. Intertegular distance 3 mm. Metasoma length 6.1 mm. Body generally metallic cupreous except as follows: mandibular apex black; head just above lower tangent of compound eyes to vertex dark, metallic green; gena dark, metallic green; apices of claws black; forewing somewhat infuscate on anterior third beyond stigma, otherwise very weakly infumate. Head narrower than mesoscutum, dorsally about 3.3 times as broad as long; occipital carina bordering lateral ocelli posteriorly, sharp, running down till just below centre of compound eyes. Scape minutely punctured, punctures separated by about a puncture width, integument between smooth. Upper carinate margin of scrobes less than one-half diameter of median ocellus from median ocellus, sharp along all margins (figure 3). Face with small, contiguous punctures; punctures slightly larger on vertex. Interantennal area weakly convex, with weak median carina. Gena punctured as on face. Head with exceedingly sparse, minute setae. Pronotum, mesoscutum and scutellum with coarse, regular punctures, punctures contiguous (figure 6); those along posterior margin of scutellum elongate posteriorly, approximately two to three times longer than wide. Pronotum with strong, premarginal carina. Posterior margin of scutellum slightly depressed and forming a short lamella that slightly overhangs anterior margin of dorsellum. Dorsellum with two rows of strong alveolae, integument within smooth, lateral and posterior margins strongly carinate. Metanotum with single row of alveolae along posterior margin, anterior half with small, contiguous punctures. Propodeum two times wider than dorsellum, with coarse, contiguous punctures, medial carina absent. Mesopleuron above depression with coarse, contiguous punctures (figure 2); depression and preepisternum smooth with sparse, minute punctures. Metacoxa with coarse, contiguous punctures except large impunctate area on dorsal surface (figure 5), depression in posterior-dorsal surface impunctate; lateral surface with smaller punctures than those of dorsal surface, punctures separated by width or less, integument between smooth. Metafemur not very stout; ventral margin with broad basal tooth followed by seven smaller teeth (figure 5). Metatibia apex truncate; outer metatibial spur long; metabasitarsus dorsally longer than breadth of metatibia (figure 5). Mesosoma with sparse, minute setae except propodeum without pubescence. Metasoma hardly longer than combined lengths of head and mesosoma. Anterior third of T 1 (of gaster) impunctate, middle third with coarse, contiguous punctures, punctured area bordered posteriorly by short band of impunctate integument, posterior margin with a short band of coarse, contiguous punctures; T 1 with premarginal constriction at point of posterior impunctate band. Remaining terga uniformly punctured, punctures coarse and contiguous. Ovipositor relatively short (figure 4). Metasoma with exceedingly sparse, minute setae. Male: Unknown."},{"description":"(figures 1 - 6)"},{"description":"Preservation. The specimen is magni fi cently preserved in tact and without mould (figures 1, 2, 6). There is essentially no damage to the specimen and only the position of certain body parts obscure specific views (e. g. the left forewing is positioned such that it makes viewing of the left hind leg difficult, although not impossible)."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1457":"Hymenoptera","9443":"Leucospidae","1360196":"Leucospis"},"class":"Insecta"},{"key":1360484,"nameKey":1578628,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1360484,"parentKey":1360483,"parent":"Bouceklytus","kingdom":"Animalia","phylum":"Arthropoda","order":"Hymenoptera","family":"Tetracampidae","genus":"Bouceklytus","species":"Bouceklytus arcuodens","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1457,"familyKey":3746,"genusKey":1360483,"speciesKey":1360484,"scientificName":"Bouceklytus arcuodens Yoshimoto, 1975","canonicalName":"Bouceklytus arcuodens","authorship":"Yoshimoto, 1975","publishedIn":"Yoshimoto, C.M. 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JOURBOOK: Amber and fossils VOLUME: 1(1) PAGES: 17-22,45. 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There are two occurrences in the south-central Yukon (LaForce and Ethel lakes); almost certainly the species ranges throughout the southern Yukon and probably across much of boreal Canada. Ecoprovinces and other designations. BC: Southern Interior, Southern Interior Mountains, Northern Boreal Mountains. YT: Boreal Cordillera. Range. Cordilleran (plus eastern distribution). Yukon and British Columbia east to Ontario and Quebec; Maine south to New Jersey, Pennsylvania, Ohio, Oklahoma, New Mexico, California (Webb et al. 2013). Biological notes. Flight period: 6 March – 21 July. 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Curious snipe-flies (Diptera: Rhagionidae) from the Purbeck of Dorset, the Wealden of Weald and the Lower Cretaceous of Spain and Transbaikalia. Proc. Geol. Assoc. 111: 153-160. 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Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. ZweiteTheil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. 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A specimen identified as Bibio cf. fusiformis and illustrated by Théobald (1937, Pl. XXI, Fig. 10) in FSL (FSL 391917) fits the specimens from Öhningen reasonably well but is not well enough preserved for a certain identification. Another specimen (BMNH In 39823) in BMNH is also in too poor condition to identify the species with any degree of certainty. Furthermore, there are four female specimens in MNHM (labeled with the numbers 209–212) identified as Bibio fusiformis . These share with the type specimens the conspicuously slender, cylindrical abdomen but are otherwise devoid of systematically useful characters."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","811":"Diptera","7281":"Bibionidae","1543369":"Bibio"},"class":"Insecta"},{"key":1590987,"nameKey":1460780,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1590987,"parentKey":1543369,"parent":"Bibio","acceptedKey":1590985,"accepted":"Bibio vicinus Lynch Arribálzaga, 1878","basionymKey":9378585,"basionym":"Phthiria dubia Germar, 1837","kingdom":"Animalia","phylum":"Arthropoda","order":"Diptera","family":"Bibionidae","genus":"Bibio","species":"Bibio vicinus","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":811,"familyKey":7281,"genusKey":1543369,"speciesKey":1590985,"scientificName":"Bibio dubius Bellardi, 1859","canonicalName":"Bibio dubius","authorship":"Bellardi, 1859","publishedIn":"Bellardi, L. 1859. 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Austroconops Wirth and Lee, a Lower Cretaceous genus of biting midges yet living in Western Australia: A new species, first description of the immatures and discussion of their biology and phylogeny (Diptera: Ceratopogonidae). American Museum Novitates 3449, 67 pp. [2004.08.23]","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1633111","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"TABLE 6 Data on Rearing of Austroconops annettae (2001 – 2002)"},{"description":"TYPE MATERIAL Holotype, male adult on microscope slide, labeled ‘‘ HOLOTYPE Austroconops annettae Borkent, 5 km SSW of Forest Grove, Loop Road, WA, Australia, 13 XI 2001, A. Borkent, CD 1998 ’’ (WAMP); allotype, female adult on microscope slide, labeled as for holotype and ‘‘ Female which laid eggs, resulting in 1 – 4 instar larvae’ ’ (WAMP); paratypes 3 females, 7 eggshells, 8 firstinstar larvae, 6 secondinstar larvae, 9 thirdinstar larvae, 1 fourthinstar larva, 1 terminal portion of abdomen of second or thirdinstar larva, all reared from eggs laid by female allotype; 2 females from type locality but collected 2 XI 2001 (CNCI); 1 female from Augusta, WA, 3 X 1970 (ANIC); immatures (CNCI)."},{"description":"DISTRIBUTION AND BIONOMICS Austroconops annettae is known from two sites in southwestern Australia (fig. 22 B). The type locality (fig. 22 D) is on Loop Road, 5 km SSW of Forest Grove, WA. One male and four females (one lost after being collected) were swept from a very small patch of low vegetation immediately beside a very shallow (less than 3 cm deep) small pool less than a meter in diameter (fig. 21 C). The pool was immediately beside and south of the narrow track of Loop Road and was located in a dry, shallow streambed (likely with running water during the winter). Surrounding vegetation was composed of an open Jarrah – Marri forest with thick stands of shrubs (Leucopogon parviflorus was common)."},{"description":"The distinctive claw of female A. annettae (fig. 1 J), with a pronounced inner basal tooth moreorless situated in the same plane as the rest of the claw, may indicate the type of host upon which the female feeds. Within the Simuliidae, females of species with a similarly shaped claw are restricted to those which feed on birds (Crosskey, 1990). Within the Ceratopogonidae, the only species of vertebrate feeders with variation regarding the presence or absence of an inner basal tooth are those in the genus Leptoconops (Forcipomyia Meigen (Lasiohelea Kieffer) and Culicoides all have simple claws with at most, a small, very slender spicule). Nearly all Leptoconops which bite mammals have a simple claw or a claw with a small basal spicule. Species with a claw with a pronounced inner basal tooth feed either on humans (e. g., L. spinosifrons (Carter) and L. siamensis Carter; Chanthawanich and Delfinado, 1967) or on birds (L. werneri Wirth and Atchley; Wirth and Atchley, 1973), indicating that the shape of the claw may not strictly indicate host type in Leptoconops. Other species which have a claw shape virtually identical to A. annettae are L. freeborni Wirth, L. melanderi Wirth and Atchley and L. patagoniensis Ronderos, but their hosts are unknown. Further research is warranted because we do not have host records for many species of Leptoconops. Lane"},{"description":"(1977) has shown that the claw shape of females of species of Forcipomyia (Trichohelea Goetghebuer) are adaptations to cling to the scales of butterfly wings as females feed on butterfly blood (they pierce the wing veins to obtain this). If further study shows a relationship between the claw shape of females of Austroconops and Leptoconops and vertebrate host type, it will provide the basis for interpreting such variation in the fossil record: Lebanese amber species Austroconops gladius and A. gondwanicus, 121 million years old, both have large basal teeth on their claws similar to those of A. annettae (Borkent, 2000 a). The female allotype collected on November 13 laid eggs scattered separately on the surface of wet mud in a vial or on the sides of the vial just above the mud. The sequence of egg laying, hatching, and larval development is given in table 6. Larval behavior was indistinguishable from that of A. mcmillani (see above). In addition to the feeding observations associated with the fecal infusion described under A. mcmillani, one firstinstar larva A. annettae was observed to eat a small live nematode whole (in less than 2 seconds). A second larva, upon encountering a somewhat moribund large nematode, pierced it at midlength. This was followed by rapid movement of the pharyngeal complex and ingestion of part of the nematode; it did not complete feeding on the nematode. In spite of these observations, the first to fourthinstar larvae mostly ignored the large number of nematodes added to the Petri dishes every 2 – 3 days and, similar to larvae of A. mcmil lani, congregated near fresh drops of the fecal infusion, suggesting that they too primarily feed on microorganisms."},{"description":"DIAGNOSIS: Male. The only extant Austroconops with flagellomere 12 short, 0.4 the length of flagellomere 13 (fig. 1 B). Female. The only extant Austroconops with each claw with a well developed basal tooth (fig. 1 J). Egg and larva (all instars). Not distinguishable from those of A. mcmillani (see generic diagnosis above). Pupa. Unknown."},{"description":"The single female from Augusta (fig. 22 B) was collected with a sweepnet and was either collected 1.6 km south of Augusta (most likely) or about 1.6 km east of Jewel Cave (about 7.5 km NW of Augusta) (D. Colless, personal commun.). Two females found on November 2, 2001 at the type locality were collected between 3: 00 and 4: 00 PM when the ambient temperature was 20 ° C (but substantially warmer in the sun). When the single female retained live in a vial was at 17 ° C, she became very lethargic (nearly torpid), but when the vial was warmed up, she again became very active. A third female collected on November 4 was lost but was collected at 12: 50 PM and at 22 ° C. One male was collected on November 13 at 1: 15 PM at 33 ° C, and one female was found on the same date and temperature at 3: 30 PM. Fervent and repeated sweeping all around this immediate site on each of the above dates failed to produce any further specimens, which suggests that the few adults were indeed concentrated at the small pool that appeared to be the only open water (although very small) in a large area. Samples of mud from the edge and bottom of this small pool failed to produce any immatures."},{"description":"TAXONOMIC DISCUSSION Austroconops annettae is very similar to A. mcmillani in all its stages (pupa not known) and we were unable to distinguish differences in male or female genitalia, eggs, or the different larval instars (except for some size differences in the immatures, which may have been due to restricted sample size or laboratory rearing conditions; tables 3, 4). The firstinstar larvae are described as having nine undivided segments (fig. 2 C). However, in A. mcmillani, early firstinstar larvae also appear to have undivided segments, whereas older firstinstar larvae have divided segments. The same is likely true for A. annettae."},{"description":"In addition to the above material, more eggs, eggshells, and specimens of the different larval instars were studied but were either left to develop further or were subsequently lost. Therefore, the numbers in tables 3 and 4 that record specimens do not match that listed above. DERIVATION OF SPECIFIC EPITHET This species is named after the first author’s wife, Annette Borkent. She shared in virtually every aspect of the sixweek expedition to study Austroconops, and the results of this paper would not have been possible without her continuous support."},{"description":"DESCRIPTION: Male adult: Descriptive statistics in table 1. Head: Antenna with well developed plume. Flagellomere 12 elongate, with subbasal constriction, 0.43 length of flagellomere 13 (fig. 1 B). Mouthparts moderately long. Palpus (fig. 1 D) with 4 segments, segment 3 slightly ovoid in lateral view (more slender than in A. mcmillani), with capitate sensilla scattered on surface or in shallow pits. Thorax: Scutellum angular in dorsal view. Wing (fig. 1 K): Costa extending just beyond apex of R 3. Legs: Legs lacking armature. Bristles on midleg tibia, first tarsomere elongate (fig. 1 N). Midleg tibia without apical spur. Hindleg first tarsomere without thick basal spine or stout setae. Claws simple, each claw apically bifid. Genitalia: In life, rotated about 50 °. Not distinguishable from that of A. mcmillani. Female adult: Descriptive statistics in table 2. Head: Ommatidia narrowly separated dorsomedially. Antenna as for A. mcmillani. Flagellomeres gradually increasing in length from flagellomere 2 to 13. Mouthparts moderately elon gate, mandible (fig. 1 H) narrow, with fine teeth, most directed dorsolaterally, laciniae with well developed, fine retrorse teeth. Palpus (fig. 1 F) with 4 segments, segment 3 ovoid, slightly swollen in lateral view, with capitate sensilla scattered on surface or in shallow pits. Thorax: Scutellum angular in dorsal view. Wing (fig. 1 L): Costa extending to or just beyond apex of R 3. Legs: Femora, tibiae slender. Legs lacking armature. Midleg tibia without apical spur. Hindleg first tarsomere without thick basal spine or stout setae. Foreleg, midleg, hindleg claws (fig. 1 J) nearly straight for apical threefourths, with stout basal tooth. Genitalia: Indistinguishable from that of A. mcmillani. Egg: Descriptive statistics as in table 3. Firstinstar larva: Head capsule length statistics in table 4. Total body length 0.67 – 0.77 mm (N = 7). Secondinstar larva: Head capsule length statistics in table 4. Total body length 1.43 – 1.82 mm (N = 4). Thirdinstar larva: Head capsule length statistics in table 4. Total body length 2.05 – 2.61 mm (N = 8). Fourthinstar larva: Head capsule length statistics in table 4. Total body length uncertain. Pupa: unknown."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","811":"Diptera","3340":"Ceratopogonidae","1633108":"Austroconops"},"class":"Insecta"},{"key":1633112,"nameKey":1221274,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1633112,"parentKey":1633108,"parent":"Austroconops","kingdom":"Animalia","phylum":"Arthropoda","order":"Diptera","family":"Ceratopogonidae","genus":"Austroconops","species":"Austroconops megaspinus","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":811,"familyKey":3340,"genusKey":1633108,"speciesKey":1633112,"scientificName":"Austroconops megaspinus Borkent, 2000","canonicalName":"Austroconops megaspinus","authorship":"Borkent, 2000","publishedIn":"Borkent, A. 2000. 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Fossil Psychodidae in Mexican amber, Part 2. (Diptera: Insecta). J. Paleont. 37: 110-118.","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1640057","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"† Psychoda usitata Quate, 1963 usitata Quate, 1963a: 116 (as Psychoda). Type locality: Mexico, Chiapas, Simojovel (Oligocene/Miocene). Additional references: Evenhuis (2012, fossil catalogue). General distribution. Mexico (Quate 1963a). Distribution in Mexico. 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New genera and species of psychodoid flies from the Lower Cretaceous amber of Lebanon. Palaeontology 42: 1101-1136. 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New genera and species of psychodoid flies from the Lower Cretaceous amber of Lebanon. Palaeontology 42: 1101-1136. 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Fossil Psychodidae in Mexican amber, Part 2. (Diptera: Insecta). J. Paleont. 37: 110-118.","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1641567","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"† Philosepedon labecula Quate, 1963 labecula Quate, 1963a: 116 (as Philosepedon). Type locality: Mexico, Chiapas, Simojovel (Oligocene/Miocene). Additional references: Evenhuis (2012, fossil catalogue). General distributio n. Mexico. Distribution in Mexic o. 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Additional references: Evenhuis (2012, fossil catalogue)."}],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","811":"Diptera","9164":"Psychodidae","1642208":"Brunettia"},"class":"Insecta"},{"key":1642437,"nameKey":11422353,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1642437,"parentKey":1642407,"parent":"Trichomyia","kingdom":"Animalia","phylum":"Arthropoda","order":"Diptera","family":"Psychodidae","genus":"Trichomyia","species":"Trichomyia glomerosa","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":811,"familyKey":9164,"genusKey":1642407,"speciesKey":1642437,"scientificName":"Trichomyia glomerosa Quate, 1963","canonicalName":"Trichomyia glomerosa","authorship":"Quate, 1963","publishedIn":"Quate, L.W. 1963. Fossil Psychodidae in Mexican amber, Part 2. (Diptera: Insecta). J. 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Fossil Psychodidae in Mexican amber (Diptera: Insecta). J. Paleont. 35: 949-951.","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1642479","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[{"description":"Distribution in Mexic o. Chiapas (Quate 1963 a)."},{"description":"General distributio n. 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New true flies (Insecta: Diptera) from the Lower Cretaceous of southern England. Cretaceous Res. 22: 451-460. 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[In Russian] [1993.??.??]","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1679487","extinct":true,"habitats":[],"nomenclaturalStatus":[],"threatStatuses":[],"descriptions":[],"vernacularNames":[],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","811":"Diptera","3342":"Chaoboridae","1679486":"Mesocorethra"},"class":"Insecta"},{"key":1687626,"nameKey":7395083,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1687626,"parentKey":1687617,"parent":"Neduba","kingdom":"Animalia","phylum":"Arthropoda","order":"Orthoptera","family":"Tettigoniidae","genus":"Neduba","species":"Neduba extincta","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1458,"familyKey":3991,"genusKey":1687617,"speciesKey":1687626,"scientificName":"Neduba extincta Rentz, 1977","canonicalName":"Neduba extincta","authorship":"Rentz, 1977","publishedIn":"Rentz, D.C.F. (1977) A new and apparently extinct katydid from Antioch sand dunes. Entomological News, 88, 241–245. Available from http://www.biodiversitylibrary.org/item/55166#page/245/mode/1up","nameType":"SCIENTIFIC","taxonomicStatus":"ACCEPTED","rank":"SPECIES","origin":"SOURCE","numDescendants":0,"numOccurrences":0,"taxonID":"gbif:1687626","extinct":true,"habitats":["TERRESTRIAL"],"nomenclaturalStatus":[],"threatStatuses":["EXTINCT"],"descriptions":[{"description":"Type material. The holotype male is the only specimen known. Images of the holotype are available at OSFO (Cigliano et al. 2020)."},{"description":"Distribution. Antioch Sand Dunes, Contra Costa County, California, on the western edge of the Central Valley. Habitat. Historically known from the sandy banks of the San Joaquin River, elevation 9 m. The 55 acre Antioch Dunes National Wildlife Refuge is the only National Wildlife Refuge in the country established to protect endangered plants and insects. Seasonal occurrence. The only known specimen was collected 1 - VII- 1937 (ES Ross, CAS). Stridulatory file. (n = 1) length 3.2 mm, 167 teeth, tooth density 52.2 teeth / mm, Song. Unknown. Karyotype. Unknown. Recognition. Description indicates large body size and absence of styli on subgenital plate. The stridulatory file tooth density places this species at the upper end of variation for the Sequoia Group. The tooth density of the single known specimen is less than the closest geographic relative, N. inversa (density 64 – 68 teeth / mm). The single specimen is the only Sequoia Group individual collected in the Central Valley west of the Sierra Nevada. Notes. This species is one of four extinct North American Orthoptera species (Hoekstra 1998) and that status has not changed; David Rentz and DBW have searched for this species at the type locality on several occasions over the last few decades, visiting during summer months when Neduba are active and using a bat detector. On no occasion were individuals found. The lack of molecular, bioacoustical, and cytogenetic characters make this species difficult to place in context of this revision, but the stridulatory file tooth density is consistent with the Sequoia Group. Minimally destructive molecular work may be undertaken in the future to place N. extincta into phylogenetic context. Among the extant Sequoia Group species, N. inversa is distributed near the San Joaquin River watershed and is therefore a possible relative, and this lineage could have colonized the western edge of the Central Valley across riparian corridors. The description of N. arborea in this work reports the only other Sierranus or Sequoia Group members known west of the Sierra Nevada. Many Neduba populations were no doubt extirpated as the eastern slope of the Coast Ranges became more arid and as humans modified the Central Valley for agriculture. In the case of the Antioch dunes, sand mining and commercial development drove extinction of this species. Thorough collecting efforts are needed in the eastern slopes of the Coast Ranges to search for possible unknown populations."},{"description":"Measurements. See Rentz (1977)."},{"description":"Fig. 19 (distribution). Common name. Extinct Shieldback. History of recognition. Described from a single male museum specimen deposited at CAS (Rentz 1977)."},{"description":"Terrestrial."},{"description":"Holotype."}],"vernacularNames":[{"vernacularName":"Antioch Dunes Shieldback Katydid","language":"eng"},{"vernacularName":"Extinct Shieldback","language":"eng"},{"vernacularName":"Antioch Dunes Shieldback Katydid","language":"eng"},{"vernacularName":"Antioch Dunes Shieldback Katydid","language":"eng"},{"vernacularName":"Extinct Shieldback","language":"eng"},{"vernacularName":"Antioch Dunes Shieldback Katydid","language":"eng"}],"synonym":false,"higherClassificationMap":{"1":"Animalia","54":"Arthropoda","216":"Insecta","1458":"Orthoptera","3991":"Tettigoniidae","1687617":"Neduba"},"class":"Insecta"},{"key":1699392,"nameKey":6817566,"datasetKey":"d7dddbf4-2cf0-4f39-9b2a-bb099caae36c","constituentKey":"7ddf754f-d193-4cc9-b351-99906754a03b","nubKey":1699392,"parentKey":1698834,"parent":"Melanoplus","basionymKey":9131822,"basionym":"Caloptenus spretus Walsh, 1866","kingdom":"Animalia","phylum":"Arthropoda","order":"Orthoptera","family":"Acrididae","genus":"Melanoplus","species":"Melanoplus spretus","kingdomKey":1,"phylumKey":54,"classKey":216,"orderKey":1458,"familyKey":9394,"genusKey":1698834,"speciesKey":1699392,"scientificName":"Melanoplus spretus (Walsh, 1866)","canonicalName":"Melanoplus spretus","authorship":"(Walsh, 1866) ","publishedIn":"Walsh, B.D. (1866) Grasshoppers and Locusts. Practical Entomologist, 2(1), 1–5. 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(1936) Orthoptères fossiles et subfossiles de l'ambre et du copal. Annales de la Société Entomologique de France, 105, 375–386. 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